STANFORD    UNIVERSITY    PUBLICATIONS 
UNIVERSITY    SERIES 

BIOLOGICAL    SCIENCES 

VOLUME  I  NUMBER  3 


New  Species  of  Melyridae, 
Chrysomelidae  and  Tenebrionidae 

(COLEOPTERA) 

From  the  Pacific  Coast,  with  Notes 
On  Other  Species 


FRANK  E.  BLAISDELL,  SR. 

Associate  Professor  of  Surgery 
Permanent  Secretary  of  the  Pacific  Coast  Entomological  Society 


STANFORD  UNIVERSITY,  CALIFORNIA 

PUBLISHED  BY  THE  UNIVERSITY 

1921 


STANFORD  UNIVERSITY 

Fust 


CONTENTS 

INTRODUCTION .        .        .'  137 

PART  I. 

Family  Melyridae  .        .        ...       .        .                         .        .  139 

Listrus  Mots  . .'        .        .        .  139 

Description  of  new  species       .         .         .         .         .         .         .  141 

Synopsis  of  the  species  of  Listrus   .        .        .        ...  175 

Dasytes  Payk.,  new  species  of                .         .        .         .•                 .  182 

Synopsis  of  the  species  of  Dasytes  .         .         .         .         .        .,  185 

Distribution  of  the  species  of  Dasytes,  Notes  on     ,    "   .         .         .  187 

Listrimorpha,  new  genus  and  species     ......  188 

Dasytastes  vanduzeei,  new  species 189 

Bibliography  and  cabinet  arrangement 190 

PART  II. 

Chrysomelidae 195 

Glyptoscelis,  new  species  of 195 

Tenebrionidae 198 

Centrioptera,  new  species  of 198 

Schizillus  laticeps  Horn 201 

New  species  of               203 

Euschides  lecontei  Horn 208 

Gigantea,  new  subspecies  of 209 

Coniontis  santarosae,  new  species 209 

musculus  Blais 210 

Coniontides  Casey 211,  212,  213 

vandykei,  new  species  of        .         .         .                 .         .  212 

Eusattus  difficilis  Lee 213 

vanduzeei,  new  species 214 

Eleodes,  new  species  of 217 

Eleodes  rotundipennis  Lee 217 

Tenebrio  tenebroides  Beauv. 226 

Helops,  new  species  of,  with  remarks  on  related  species        .         .  226 

Synoptic  table  of  Helops,  in  part 230 

Addendum                       231 


INTRODUCTION  137 


INTRODUCTION 

The  material  on  which  the  present  avocational  studies  are  based,  was 
collected  chiefly  in  California,  the  smaller  part  having  been  taken  in 
Washington,  Oregon,  Nevada,  Lower  California  and  Islands  off  the 
coast.  The  following  Coleoptera  are  described  as  new  to  science :  Twenty 
species  and  one  subspecies  of  Listrus',  two  species  and  one  variety  of 
Dasytes;  one  species  of  Dasytastes;  one  species  and  the  genus  Listri- 
morpha;  two  species  of  Glyptoscelis ;  two  species  of  Centrioptera;  three 
species  of  Schizillus;  one  species  of  Coniontis;  one  species  of  Coniontides; 
one  species  of  Eusattus ;  five  species  and  three  races  of  Eleodes  and  three 
species  of  Helops.  Notes  on  a  few  other  species  are  also  given.  At  all 
times  an  effort  has  been  made  to  examine  long  series,  and  uniques  have 
been  considered  only  when  they  presented  well  marked  characters,  and 
even  then  collateral  evidence  was  at  times  obtained  from  colleagues  or 
collectors  in  the  field. 

I  am  greatly  indebted  to  the  California  Academy  of  Sciences  for  the 
privilege  of  studying  the  material  in  the  collection  of  its  entomological 
department;  also,  to  Mr.  Van  Duzee,  curator  of  the  department,  for 
personal  favors.  The  following  friends  and  collectors  have  responded 
generously  to  my  requests  for  material  and,  to  them  I  express  my  sincere 
thanks:  Mr.  J.  O.  Martin  for  the  liberal  gift  of  specimens  from  the 
Bear  Lake  region  in  San  Bernardino  County,  and  Mr.  F.  W.  Nunen- 
macher  for  similar  gifts  and  the  opportunity  of  studying  long  series  of 
specimens  from  different  regions  in  California.  Mr.  Chas.  Liebeck  of 
Philadelphia  very  kindly  loaned  the  material  in  his  collection.  To  Mr. 
H.  C.  Fall,  Mr.  Chas.  L.  Fox  and  Mr.  O.  N.  Sanford  my  appreciation  is 
due  for  less  extensive  but  none  the  less  valuable  assistance. 


FAMILY    MELYRIDAE  139 


PART    I. 

Family  Melyridae. 

As  recently  decreed  by  the  leading  Coleopterists  of  the  United  States, 
the  old  family  name  Malachidae  gives  way  to  Melyridae1;  the  latter 
includes  three  subfamilies,  namely:  Malachiinae,  Melyrinae  and  Rhada- 
linae. 

Nothing  of  any  extent  has  been  written  on  the  American  Melyrinae 
since  Col.  Casey's  excellent  paper  published  in  1895  2.  Fall  added  a  few 
species  in  1901  3  and  1907 4.  In  1906 6  I  described  two  new  species  of 
Dasytes. 

Listrus  Mots. 

The  genus  Listrus  is  proving  to  be  more  extensive  and  a  less  homo- 
geneous aggregate  than  was  at  first  suspected.  Its  specific  units  are 
characterized  by  an  elongate  and  more  or  less  subparallel  convex  body, 
more  or  less  sparse  and  variegated  vestiture  without  intermixed  setae, 
tarsi  more  or  less  variable  as  to  length  and  stoutness,  with  the  basal  joint 
of  the  posterior  slightly  longer  than  the  second,  the  fifth  somewhat 
dilated  toward  the  tip  and  canaliculate  above  at  apex  as  usual  throughout 
the  family,  the  anterior  tibiae  slender,  cylindrical,  usually  with  closely 
decumbent  ashy  pubescence  and  only  occasionally  with  two  or  three  very 
slender,  distant,  external  spines,  these  being  completely  obsolete  as  a  rule. 

Antennae  somewhat  variable  as  to  length,  more  or  less  feebly  incras- 
sate,  only  slightly  though  distinctly  serrate,  with  the  first  joint  always 
much  dilated  and  stout  apically  and  usually  darker  in  color,  the  fifth 
larger  than  the  fourth  or  sixth,  usually  triangular,  sometimes  the  sixth 
and  eighth  joints  are  smaller  than  the  seventh,  and  the  eleventh  more  or 
less  evenly  elongate-ovoidal  or  obovate  and  obtusely  acuminate  or  truncate 
at  tip.  The  pronotum  is  dilated  behind  the  middle,  with  the  lateral  edge 
more  or  less  minutely  serrulate  and  having  an  even  fringe  of  short 
superiorly  recurved  setae,  which  are  almost  invariably  cinereous  in  color, 
the  apical  angles  usually  more  or  less  rounded.  Epipleura  very  narrow 
but  dilated  toward  base,  with  their  plane  strongly  inclined  upward 
throughout.  The  ungual  appendages  are  thick,  equal  and  fully  as  long 


1  "Leng's  Catalogue  of  the  Coleoptera  of  America  North  of  Mexico,"  145,  1921. 

*  "Coleopterological  Notices,"  6,  Annals  New  York  Academy  of  Sciences,  8, 
456,  July,  1895. 

8  "Occasional  Papers,"  California  Academy  of  Sciences,  8,  248-251,  1901. 

4  "The  Coleoptera  of  New  Mexico,"  Transactions  of  the  American  Entomo- 
logical Society,  33,  236-240,  1907  (Malachidae). 

'Entomological  News,  17,  74-76,  March,  1906. 


140      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

as  the  claws,  the  apical  third  or  fourth  of  which  is  free.  The  above 
characterization  has  been  modified  from  Casey. 

Fall  has  called  attention  to  the  fact  that  the  males  of  all  species  have 
the  protibiae  mucronate  at  the  tip.  The  spurs  of  the  meso-  and  meta- 
tibiae  are  variable,  especially  on  the  latter ;  generally  moderately  short  and 
blunt  on  the  former.  A  study  of  the  spurs  and  mucros  will  constitute  a 
special  line  of  investigation.  A  monographic  treatment  of  the  genus 
should  not  be  attempted  until  a  greater  amount  of  collecting  has  been 
done  and  a  larger  series  of  a  greater  number  of  species  been  secured. 

From  my  own  studies  it  is  evident  that  for  the  proper  description  and 
diagnosis  of  the  species,  attention  must  be  paid  to  the  structure  of  the 
antennae,  pronotum,  pattern  of  maculation,  fifth  ventral  abdominal  seg- 
ment, and  the  legs,  especially  the  femora  and  the  metatrochanters. 

I  have  assumed  that  the  fundamental  type  of  maculation  in  its  discrete 
form,  consisted  of  four  transverse  fasciae  of  more  or  less  cinereous 
pubescence  that  divided  the  dark  elytral  field  into  five  dark  fasciae  which 
may  be  known  as  the  basal,  post-basal,  median  or  submedian,  subapical 
and  apical ;  it  is  by  the  extension  of  the  light  fasciae  that  the  dark  areas  are 
broken  up,  or  by  the  diffusion  and  coalescence  of  the  dark  fasciae  or 
maculae  that  the  varied  pattern  of  maculation  is  produced.  This  con- 
ception of  a  definite  fundamental  pattern  of  maculation  will  help  greatly 
in  the  interpretation  of  the  species  from  color  markings. 

It  is  necessary  to  caution  students  who  endeavor  to  identify  the 
different  species,  that  it  is  absolutely  essential  for  the  proper  recognition 
and  appreciation  of  antennal  characters  to  view  the  several  joints  at 
right  angles  to  their  broad  surfaces,  which  according  to  my  terminology 
are  the  dorsal  and  ventral  surfaces;  viewing  the  antennae  edgewise — 
antero-posteriorly — determines  the  degree  of  compression.  The  care- 
less viewing  of  the  surface  at  a  tangent  will  result  in  an  erroneous  con- 
ception of  the  relative  proportions  of  the  parts. 

It  is  not  sufficient  to  simply  examine  the  specimens  with  a  low-power 
hand  lens,  but  with  as  powerful  a  glass  as  can  be  obtained.  The  failure 
to  recognize  the  many  undescribed  species  in  the  past  has  been  due  to 
the  inability  to  see  the  abundance  of  good  specific  characters.  In  the 
present  study  the  binocular  and  stereoscopic  microscopes  were  used. 

DESCRIPTIONS  OF  NEW  SPECIES. 

Listrus  cephalicus,  new  species. — Form  oblong-oval,  very  moderately 
convex.  Color  black,  upper  surface  with  a  bluish  or  subaeneous  metallic 
lustre;  antennae  and  legs  nigro-piceous. 

Pubescence  moderately  short  and  sparse,  recumbent,  cinereous  hairs 
not  abundant;  brownish-black  hairs  arranged  in  an  indefinite  pattern  in 
which  the  maculae  and  fasciae  observed  in  other  species  diffuse  and 


DESCRIPTION  OF  NEW  SPECIES  141 

coalesce,  so  that  the  prevailing  color  is  dark.  Pronotal  maculae  not  dis- 
cernible on  account  of  the  abundance  of  dark  hairs. 

Head  comparatively  large  and  moderately  transverse ;  eyes  large  and 
more  strongly  convex  in  anterior  two-thirds  and,  there  unusually  promi- 
nent, anterior  surface  quite  abrupt,  posteriorly  forming  a  straight  line 
with  the  tempora  which  are  quite  strongly  convergent  behind;  frons 
broad,  rather  strongly  and  very  broadly  impressed,  surface  densely 
indentato-punctate,  punctures  coarser  and  less  crowded  on  the  vertex. 
Antennae  long  and  stout,  slightly  compressed,  gradually  but  not  strongly 
incrassate;  second  joint  stout,  as  wide  as  the  sixth,  nearly  oval  but 
slightly  narrowed  at  base;  third  subequal  in  length  to  the  second,  rather 
strongly  obconical  and  almost  twice  as  long  as  wide ;  fourth  subtriangular, 
about  as  long  as  wide,  subserrate  anteriorly,  apical  margin  oblique  and 
about  as  wide  as  the  seventh ;  fifth  subequilaterally  triangular,  distinctly 
serrate  anteriorly,  apical  margin  oblique  and  about  as  wide  as  the  ninth ; 
sixth,  seventh  and  eighth  subequal  in  size,  sixth  triangular,  about  as  long 
as  wide,  subserrate  anteriorly;  seventh  very  slightly  longer,  triangular; 
eighth  slightly  transverse  and  triangular;  ninth  stouter,  wider  than  long, 
anterior  margin  not  strongly  arcuate ;  tenth  wider  than  long,  sub- 
triangular  with  sides  distinctly  arcuate ;  eleventh  obovate,  not  wider  than 
the  tenth,  truncate  at  tip,  almost  a  half  longer  than  wide. 

Pronotum  almost  a  third  wider  than  long,  widest  just  behind  the 
middle;  apex  arcuato-truncate  in  circular  arc;  apical  angles  quite 
broadly  rounded;  sides  rather  strongly  and  evenly  arcuate  in  posterior 
two-thirds,  thence  straight  and  rather  strongly  convergent  to  apex, 
serrules  strong,  subobtuse,  about  as  long  as  wide  at  base,  fimbria  rather 
short  and  not  closely  placed ;  base  quite  broadly  and  moderately  arcuate, 
rather  broadly  but  not  strongly  sinuate  within  the  basal  angles,  which  are 
more  or  less  obtuse;  disk  broadly  and  quite  evenly  convex  posteriorly, 
more  strongly  so  anteriorly  and  arcuately  declivous  antero-laterally, 
densely  and  rather  strongly  indentato-punctate,  punctures  rather  coarse 
and  strong. 

Elytra  oblong,  rather  less  than  twice  as  long  as  wide,  humeri  some- 
what prominent,  base  rather  transverse;  disk  quite  coarsely  punctate, 
surface  more  or  less  transversely  rugose  or  wavy,  smoother  toward  apex 
where  the  punctures  are  finer  and  not  impressed;  feeble  parascutellar 
prominences  rather  discretely  punctured  and  not  rugose.  Scutellum 
quadrate. 

Abdomen  finely  punctulate  and  densely  microreticulate ;  broad  at 
base,  sides  converging  rather  strongly  toward  apex. 

Legs  rather  stout  and  somewhat  long ;  tarsi  elongate  and  also  stout. 

Male.— Oblong-suboval.     Metafemora    broad    and    moderately    flat- 


142      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

tened;  metatibiae  distinctly  thickened  in  apical  two-thirds,  very  feebly 
arcuate;  fifth  ventral  abdominal  segment  about  as  long  as  the  second, 
broadly  and  transversely  truncate  at  apex ;  a  sixth  segment  is  visible  and 
the  under  surface  of  the  pygidium  rather  flat  and  fringed  with  quite  short 
and  stout  black  hairs. 

Female  unknown. 

Measurements. — Length,  3.4  mm.;  width,  1.2  mm.  Holotype,  male, 
in  the  collection  of  the  California  Academy  of  Sciences.  Described  from 
a  unique,  collected  by  Mr.  E.  P.  Van  Duzee,  on  April  21st,  1918. 

Type  locality. — Salada  Beach,  San  Mateo  County,  California. 

Cephalicus  is  a  very  distinct  and  unique  species  in  its  prominent  and 
peculiarly  convex  eyes,  broad  head,  stout  antennae  and  legs,  besides  the 
preponderance  of  dark  hairs  on  the  upper  surface  with  consequent 
obscuration  of  the  maculation. 

Listrus  coloradensis,  new  species. — Form  oblong,  subovate,  moder- 
ately convex.  Color  black,  surface  rather  dull  and  with  a  feeble  bluish 
lustre ;  antennae  piceous,  legs  rufo-piceous. 

Pubescence  very  short,  rather  coarse  and  more  or  less  squamiform, 
not  dense,  rather  inconspicuous,  plumbeo-cinereous  in  color  above  but 
more  cinereous  beneath  where  it  is  somewhat  longer  and  closely  recum- 
bent ;  noticeably  coarse  on  front  of  the  head. 

Head  slightly  transverse,  eyes  rather  large  and  moderately  promi- 
ment ;  front  broadly  impressed,  more  strongly  and  somewhat  bi-impressed 
between  the  antennae,  densely  but  not  coarsely  indentato-punctate,  some- 
times with  a  small  subglabrous  convexity  close  to  the  middle  of  the 
epistomal  base.  Antennae  rather  short  and  somewhat  stout  and  nearly 
similar  in  the  sexes;  second  joint  slightly  oblong-oval,  just  a  little  longer 
than  wide ;  third  distinctly  obconical  and  nearly  twice  as  long  as  wide ; 
fourth  a  little  stouter,  subobconico-triangular  and  about  as  long  as  the 
sixth ;  fifth  subequal  to  the  ninth  in  width,  subtriangular  and  serrate,  very 
little  longer  than  wide  and  obliquely  truncate  at  apex ;  sixth  and  eighth 
subequal,  quite  distinctly  smaller  than  the  contiguous  joints,  very  slightly 
wider  than  long  and  feebly  triangular ;  seventh  triangular  and  about  as 
long  as  wide,  more  prominent  anteriorly  than  posteriorly,  slightly  longer 
than  the  sixth  or  seventh ;  ninth  and  tenth  stouter,  a  little  wider  than  long 
subtriangular,  sides  feebly  arcuate ;  eleventh  obovate-triangular,  scarcely 
a  half  longer  than  wide. 

Pronotum  about  a  fourth  wider  than  long,  quite  equal  in  width  to  the 
elytral  base;  apex  arcuato-truncate  in  circular  arc;  sides  moderately 
rounded  in  basal  half,  thence  almost  straight  to  the  apical  angles  as  viewed 
from  above,  serrules  small,  fimbriae  rather  short  and  stout ;  apical  angles 
obtusely  rounded ;  base  broadly  arcuate,  rounding  evenly  into  the  sides ; 


DESCRIPTION  OF  NEW   SPECIES  143 

disk    moderately    convex    basally,    arcuately    declivous    antero-laterally. 
densely  but  not  coarsely  indentato-punctate. 

Elytra  oblong-oval,  about  twice  as  long  as  wide,  humeri  rather 
prominent ;  disk  not  strongly  punctate,  punctures  shallow,  separated  by  a 
distance  equal  to  two  or  three  times  their  diameters,  surface  micro- 
scopically reticulate;  apex  parabolically  rounded,  sutural  angles  rather 
more  than  narrowly  rounded. 

Abdomen  finely  and  sparsely  punctate,  surface  microscopically  reticu- 
late. 

Legs  rather  short. 

Male. — Slightly  narrower.  Fifth  ventral  segment  arcuato-truncate 
at  apex.  Antennal  joints  five,  six  and  seven  similar  in  form,  subtriangu- 
lar,  more  prominent  anteriorly  than  posteriorly,  eighth  smaller  by  one- 
eighth. 

Female. — Slightly  broader.  Fifth  ventral  rather  parabolically 
rounded.  Antennae  slightly  more  slender,  joints  six  and  eight  smaller, 
similar  in  form ;  tenth  and  eleventh  feebly  transverse,  sides  arcuate  and 
symmetrical. 

Measurements.— Length  (Types),  2.9-3  mm.;  width,  1-1.2  mm. 
Holotype,  male,  and  allotype,  female,  and  two  paratypes  in  my  own 
collection. 

Type  locality. — Colorado,  without  definite  locality. 

In  senilis  the  pubescence  is  coarse,  dense  and  cinereous,  and  the  tenth 
antennal  joint  is  transverse ;  in  davicornis  the  pubescence  is  moderately 
long,  not  so  coarse,  and  the  tenth  antennal  joint  is  transverse  as  in  senilis, 
while  in  coloradensis  the  pubescence  is  coarse  on  the  head,  pronotum  and 
elytral  base,  the  antennae  are  almost  similar  in  the  sexes,  and  the  fifth 
ventral  abdominal  segment  is  arcuato-truncate  in  the  male.  Col.  Casey 
has  examined  specimens  of  this  species  and  pronounced  it  new. 

Listrus  dilutus,  new  species. — Sexes  similar  in  form,  oblong-oval, 
subparallel  and  somewhat  robust,  moderately  convex.  Color  black,  with 
a  dark  virido-metallic  lustre ;  tarsi  rufous  to  rufo-piceous. 

Pubescence  very  short  and  inconspicuous,  rather  plumbeo-cinereous 
in  color,  recumbent  and  moderate  in  abundance.  The  maculation  is 
difficult  of  determination,  dark  areas  invested  with  dark-brownish  hairs. 
Basal  fascia  is  represented  by  a  large  parascutellar  and  a  humeral  macula 
on  each  elytron ;  a  variable  post-basal  macula  at  middle  of  each  which  is 
frequently  connected  to  the  humeral  forming  a  lunule;  median  fascia 
rather  broad,  edges  feebly  zig-zag,  usually  more  or  less  interrupted  at 
suture ;  subapical  fascia  rather  wide,  more  or  less  entire  and  tending  to 
dilate  on  the  suture,  edges  rather  irregular ;  apical  maculae  variable  as  to 


144      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

size.  Pronotal  central  macula  evidently  hourglass-shaped  or  oval,  it  may 
have  a  narrow  median  reentrant  pale  line  from  apical  and  basal  margins ; 
lateral  semilunar  lines  rather  obscure;  hairs  darkish  on  front  of  head. 
Pronotal  fimbriae  moderately  short  and  pale  in  color. 

Head  somewhat  transverse,  front  nearly  plane,  feebly  and  longi- 
tudinally bi-impressed  anteriorly,  impressions  separated  by  a  feeble  con- 
vexity, which  is  more  or  less  glabrous  at  the  epistomal  base,  punctures 
moderate,  well  defined,  feebly  impressed,  separated  by  a  distance  equal 
to  one  or  two  times  their  diameter,  intervals  flat,  feebly  indentated  toward 
the  periphery;  muzzle  short.  Antennae  somewhat  slender  and  rather 
long,  very  feebly  incrassate  and  slightly  compressed,  less  noticeably  so  in 
the  outer  joints,  somewhat  dissimilar  in  the  sexes ;  first  joint  very  stout ; 
second  oval,  about  a  fourth  longer  than  wide,  narrowing  slightly  toward 
base;  third  subcylindrical,  about  twice  as  long  as  wide,  as  long  as  the 
fourth ;  fourth  moderately  compressed,  almost  subtriangular,  feebly  sub- 
angulate  anteriorly ;  fifth  elongate ;  sixth  and  seventh  equal  in  length,  less 
than  a  half  longer  than  wide,  scarcely  subtriangular ;  eighth  a  little  longer 
than  wide,  shorter  than  the  ninth,  slightly  oblong-triangular;  ninth  and 
tenth  distinctly  longer  than  wide,  somewhat  oblong-triangular;  eleventh 
elongate  oval,  slightly  narrowed  at  apex,  twice  as  long  as  wide ;  outer  joints 
not  noticeably  wider. 

Pronotum  about  a  fourth  wider  than  long,  rather  evenly  convex,  but 
more  declivous  antero-laterally ;  apex  arcuato-truncate ;  apical  angles 
broadly  rounded  into  the  sides ;  base  broadly  and  rather  strongly  arcuate, 
somewhat  lobed,  more  or  less  sinuate  laterally;  basal  angles  obtuse  and 
more  or  less  evident  but  not  strong;  sides  broadly  and  moderately 
strongly  arcuate,  straighter  and  moderately  convergent  anteriorly, 
serrules  short,  blunt  to  subacute,  intervals  equal  to  their  base ;  disk  rather 
discretely  punctured  in  middle  third,  punctures  distinct,  separated  by  a 
distance  equal  to  one  or  two  times  their  diameter,  intervals  more  or  less 
glabrous  centrally,  more  or  less  indentated  apically  and  basally,  laterally 
very  densely  indentato-reticulato-punctate,  surface  rather  dull. 

Elytra  oblong,  about  twice  as  long  as  wide,  sides  parallel,  very  feebly 
arcuate,  apex  broadly  rounded,  punctures  rather  coarse,  separated  by  a 
distance  equal  to  half  to  three  times  their  own  diameter,  surface  more  or 
less  irregular  from  feeble  transverse  impressions,  punctures  finer  and 
more  widely  spaced  toward  apex. 

Abdomen  very  finely  and  evenly  punctured,  more  densely  so  on  the 
fifth  segment  which  is  unmodified  in  the  male. 

Legs  moderate  in  length  and  stoutness ;  femora  rather  stout,  meta- 
femora  somewhat  rapidly  narrowed  at  the  trochanters ;  tarsi  rather  long 
and  slender. 


DESCRIPTION  OF  NEW  SPECIES  145 

Male.— Usually  smaller.  Fifth  antennal  joint  elongate,  oblong- 
triangular,  broadly  arcuate  anteriorly,  not  noticeably  wider  than  the  fol- 
lowing joints.  Fifth  ventral  abdominal  segment  rather  broadly  sinuato- 
truncate. 

Female.— Usually  larger.  Fifth  antennal  joint  shorter,  more  tri- 
angular and  more  subangulate  anteriorly ;  all  the  joints  rather  less  robust. 
Fifth  ventral  broadly  rounded  at  apex.  Femora  less  stout  and  more 
parallel. 

Measurements.— Length  (Types),  3-3.3  mm.;  width,  1-1.2  mm. 

Holotype,  male,  in  the  collection  of  the  California  Academy  of 
Sciences ;  allotype,  female,  and  paratypes  in  that  of  the  author. 

Type  locality. — Blood's  Meadow,  Alpine  County,  California;  col- 
lected by  the  author  on  July  14th,  1907,  at  an  elevation  of  7000  feet. 
Beaten  from  the  blossoms  of  a  white  Ceanothus,  in  company  with  Amphi- 
chroum  pallidum  Casey  and  another  species  of  Listrus  which  is  referred 
to  montanus  Casey.  Dilutus  is  distinct  in  the  similarity  of  the  sexes, 
short  dark  pubescence  and  inconspicuous  maculation.  In  montanus  the 
pubescence  and  maculation  are  well  developed,  hairs  rather  long,  form 
different  in  the  sexes  and  more  oval,  and  besides  the  antennae  are  dis- 
tinctly incrassate,  the  eleventh  joint  is  shorter,  more  robust,  oval  and 
wider  than  the  preceding  joints. 

Listrus  martini,  new  species. — Form  somewhat  elongate,  more  or  less 
oblong-ovate,  distinctly  more  than  twice  as  long  as  wide.  Color  black, 
with  a  feeble  purpureo-cupreous  lustre,  surface  shining;  mouth-parts 
pale  rufo-testaceous,  last  two  joints  of  the  maxillary  palpi  blackish ;  first 
joint  of  the  antennae  black,  second  rufo-testaceous,  succeeding  joints 
similarly  pale  becoming  gradually  infuscate  to  tip;  legs  pale  rufo- 
testaceous,  femora  rarely  slightly  infuscate. 

Pubescence  not  dense,  rather  short  and  slender,  more  or  less  plumbeo- 
cinereous,  with  broad  fasciae  of  nigro-f uscous  hairs  arranged  as  follows : 
A  moderately  wide  transverse  basal  fascia,  usually  unbroken;  post-basal 
fascia  absent ;  a  broad  median  fascia  extending  from  margin  to  margin ;  a 
similar  subapical  fascia  joining  the  median  fascia  along  the  suture; 
apical  maculae  absent.  The  median  and  subapical  fasciae  may  by  diffu- 
sion nearly  coalesce,  in  such  cases  separated  laterally  by  a  narrow  line  of 
pale  hairs,  rarely  distal  two-thirds  of  the  elytra  are  nearly  dark.  Prono- 
tal  lateral  vittae  not  distinct;  median  dark  area  oblong-oval,  extending 
from  apical  to  basal  margins,  usually  widest  at  middle  and  not  con- 
stricted as  in  the  hourglass  figure.  Head  slightly  maculate  in  the  central 
area. 

Head  rather  broad  and  relatively  short,  muzzle  small ;  front  broadly 
and  feebly  convex,  sometimes  feebly  impressed  behind  the  epistoma,  with 


146      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

a  slight  convexity  at  middle  against  the  frontal  suture ;  punctures  rather 
small  and  sparse,  slightly  denser  at  the  periphery ;  eyes  relatively  large, 
rather  finely  faceted,  and  moderately  prominent.  Antennae  rather  slen- 
der, last  three  joints  rather  heavy  and  not  compressed ;  third  joint  rather 
slender  and  subcylindrical,  fourth  and  fifth  subangulate  anteriorly,  fifth 
slightly  wider  than  the  following  three  joints ;  sixth,  seventh  and  eighth 
subequal ;  ninth  and  tenth  about  as  long  as  wide  and  thicker ;  eleventh 
nearly  as  long  as  the  preceding  two  taken  together  and  pointed  obovate. 

Pronotum  about  a  fourth  wider  than  long,  evenly  and  rather  strongly 
convex  from  side  to  side,  very  feebly  so  antero-posteriorly ;  apex  very 
feebly  arcuate  in  circular  arc,  angles  obtusely  rounded;  sides  feebly 
arcuate,  moderately  converging  toward  apex,  almost  broadly  and  feebly 
sinuate  adjacent  to  the  angles,  serrules  small  and  acute,  fimbriae  moder- 
ately short,  even  and  not  dense ;  basal  angles  obtuse  and  rather  distinct ; 
base  broadly  and  slightly  arcuate ;  disk  sparsely  punctate  in  central  third, 
punctures  moderately  small,  separated  by  at  least  one  or  three  times  their 
diameter,  intervals  flat  and  smooth,  rather  strongly  indentato-punctate  in 
lateral  thirds. 

Elytra  oblong,  rather  less  than  twice  as  long  as  wide ;  sides  feebly 
arcuate  and  parallel,  rather  broadly  rounded  at  apex;  punctures  rather 
sparse,  at  base  equal  in  size  to  those  of  the  pronotum,  finer  apically,  sur- 
face slightly  and  sparsely  rugose. 

Abdomen  finely  and  sparsely  punctate,  surface  rather  shining.  Fifth 
ventral  segment  unmodified  in  the  male. 

Legs  moderate  in  length ;  metafemora  not  in  the  least  constricted 
behind  and  adjacent  to  the  trochanters. 

Male. — Narrower  and  oblong.  Fifth  ventral  segment  truncate  at 
apex. 

Female. — Slightly  broader,  more  oblong-oval.  Fifth  ventral  sub- 
angulate at  apex  and  obliquely  arcuate  laterally. 

Measurements. — Length  (Types),  2.3-2.4  mm.;  width,  .9-1  mm. 

Holotype,  male,  in  the  author's  collection. 

Allotype,  female,  in  the  collection  of  the  California  Academy  of 
Sciences. 

Paratypes  in  both  collections  and  in  that  of  Mr.  J.  O.  Martin,  who 
collected  the  species  on  April  13th,  1917. 

Type  locality. — Mecca,  Colorado  Desert,  California. 

Distribution.— r Besides  the  specimens  from  the  type  locality,  others  in 
the  Liebeck  collection  have  been  studied,  these  were  collected  in  Utah. 
They  agree  in  every  particular  with  the  types.  A  series  of  about  thirty 
specimens  have  been  studied. 


DESCRIPTION  OF  NEW  SPECIES  147 

Martini  belongs  to  the  same  group  of  species  as  annulatus,  rubripes 
and  parvicejts  described  below.  In  rubripes  the  antennae  are  stouter, 
surface  lustre  duller,  and  the  pronotal  central  field  is  distinctly  but  not 
densely  indentato-punctate  and  the  sides  are  more  strongly  arcuate,  it  is 
found  in  Colorado.  In  annulatus  the  pronotal  field  is  indentato-punctate 
and  the  antennae  are  dark.  In  both  rubripes  and  annulatus  the  elytral 
dark  fasciae  are  distinctly  defined  and  do  not  tend  to  diffusion,  as  in 
martini  and  parvicollis.  In  parvicollis  the  pronotum  is  noticeably  small. 
My  series  of  annulatus  are  from  the  high  Sierras,  7000  feet,  and  Mono 
Lake,  Mono  County,  California. 

Listrus  maculosus  Casey. — Very  distinct  from  amplicollis  Casey  with 
which  it  is  usually  confused.  In  amplicollis  the  fifth  ventral  abdominal 
segment  is  modified  on  the  disk,  while  in  maculosus  it  is  simple ;  in  the 
latter  the  third,  fourth  and  fifth  antennal  joints  are  distinctly  compressed, 
large  and  triangular  in  form,  the  third  joint  is  furthermore  elongate  and 
comparatively  large  and  bears  a  distinct  stout  chitinous  seta  at  apical 
border  anteriorly  in  the  male ;  the  antennae  have  their  usual  form  in  the 
female. 

Maculosus  is  taken  abundantly  about  San  Francisco  and  in  Marin 
County.  I  took  a  very  large  series  in  Humboldt  County,  at  Green  Point 
Ranch  and  Willow  Creek,  in  1917.  Casey  mentions  only  having  had  a 
single  male  at  the  time  of  drawing  up  his  description. 

Listrus  vestitus,  new  species. — Form  elongate  oval,  distinctly  more 
than  twice  as  long  as  wide  and  moderately  convex.  Color  black,  surface 
more  or  less  shining ;  second,  third  and  fourth  joints  of  the  antennae  pale, 
second  always  so,  remaining  joints  more  or  less  rufo-piceous,  terminal 
ones  usually  quite  black;  mouth  and  basal  joints  of  the  palpi  pale, 
epistoma  more  or  less  so ;  tarsi  and  distal  half  or  two-thirds  of  the  tibiae 
pale.  A  metallic  lustre  scarcely  discernible,  although  the  pronotum  may 
be  slightly  subcupreous. 

Pubescence  distinctly  cinereous,  moderately  long  and  recumbent. 
Maculae  of  dark  brownish  or  blackish  hairs  are  arranged  on  the  elytra 
as  follows :  Basal  fascia  broken  into  a  smaller  humeral  and  a  larger  sub- 
scutellar  macula  on  each  elytron ;  post-basal  fascia  represented  by  a  mid- 
elytral  subquadrate  macula,  which  rarely  extends  arcuately  forward  to 
the  humeral  forming  a  lunule;  median  fascia  more  or  less  narrow,  dis- 
tinctly and  more  or  less  sharply  zig-zag,  usually  almost  interrupted  at  the 
suture  and  on  middle  of  each  elytron;  subapical  fascia  broader,  usually 
interrupted  at  middle  of  each  elytron,  the  median  portions  forming  a  more 
or  less  rhomboidal  and  usually  rather  large  macula  across  the  suture; 
apical  maculae  variable  in  size  and  form.  Pronotal  central  macula  more 


148      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

or  less  constricted  at  middle,  with  reentering  slender  median  lines  of 
paler  hairs  at  apex  and  base ;  lateral  lunules  more  or  less  distinct. 

Head  wider  than  the  pronotal  apex;  frons  more  or  less  broadly 
impressed  and  usually  a  small  glabrous  area  is  present  behind  the  base  of 
the  epistoma,  punctures  distinct  and  discrete,  intervals  somewhat 
indentato-punctate,  surface  rugulose  at  the  epistoma.  Eyes  rather  large 
and  moderately  strongly  prominent.  Antennae  moderately  long  and 
different  in  the  sexes. 

Pronotum  about  a  fourth  wider  than  long,  moderately  and  evenly 
convex,  more  strongly  so  anteriorly  and  laterally;  apex  arcuato-truncate 
and  distinctly  narrower  than  the  base;  apical  angles  rather  broadly 
rounded ;  base  broadly  and  rather  strongly  arcuate,  angles  not  evident  and 
almost  continuously  rounded  with  the  broadly  arcuate  sides,  which  become 
less  arcuate  and  moderately  convergent  to  apex,  serrations  small  and 
rather  acute ;  disk  rather  discretely  punctured  in  the  central  area,  intervals 
flat  and  as  wide  or  wider  than  the  punctures,  almost  feebly  indentate  in 
part,  lateral  thirds  densely  rugoso-indentato-punctate. 

Elytra  oblong-oval,  punctures  rather  smaller  than  on  the  pronotum, 
and  separated  by  a  distance  equal  to  two  times  their  diameter,  somewhat 
smaller  on  apical  third ;  surface  more  or  less  feebly  transversely  impresso- 
rugose ;  apex  rather  broadly  rounded. 

Abdomen  quite  densely  punctato-rugulose ;  fifth  ventral  abdominal 
segment  not  modified  on  the  disk. 

Legs  moderate  in  length  and  stoutness. 

Male. — Narrower  elongate  oval.  Smaller  males  not  distinctly 
parallel.  Antennae  rather  stout,  outer  three  joints  incrassate  and  not 
compressed;  first  joint  about  as  long  as  wide,  second  a  little  longer  than 
wide;  third  obconico-cylindrical,  equal  to  the  fourth  in  length;  fourth 
subtriangular,  obtusely  angulate  anteriorly,  wider  than  the  third;  fifth 
a  little  longer  than  wide,  triangular,  distinctly  wider  than  the  next  three 
following  joints ;  sixth  subtriangular,  equal  to  the  seventh  in  length ; 
seventh  less  subtriangular;  eighth  less  compressed  than  the  preceding 
joints,  a  little  longer  than  wide,  more  obconic  than  triangular ;  ninth, 
tenth  and  eleventh  joints  more  robust;  ninth  as  long  as  wide,  sides 
oblique;  tenth  as  long  as  wide  and  subquadrate  in  outline,  circular  in 
transverse  section ;  eleventh  wider  than  preceding  joints,  obtusely  obovate 
and  about  a  half  longer  than  wide.  Fifth  ventral  segment  rather  broadly 
and  feebly  sinuate  at  apex;  surface  rather  asperately  sculptured,  hairs 
scarcely  darker. 

Female. — Somewhat  broader.  Antennae  more  slender,  relative 
length  and  size  of  the  joints  similar;  second  and  third  rather  elongate 
and  equal  in  length,  sixth  not  noticeably  smaller  than  the  seventh. 


DESCRIPTION  OF  NEW   SPECIES  149 

Measurements. — Length  (Types),  3.2-3 .4  mm.;  width,  1.1-1.3  mm. 

Holotype,  male,  allotype,  female,  in  my  own  collection;  paratypes 
in  that  of  Mr.  J.  O.  Martin,  who  collected  them  pn  June  5th,  1919. 

Type  locality. — Bear  Lake,  San  Bernardino  County,  California. 
Eight  specimens  studied — four  of  each  sex. 

Vestitus  is  quite  distinct  from  elegantulus  and  bifasciatus,  where  the 
males  are  distinctly  elongate  and  parallel.  In  vestitus  the  differences  in 
form  between  the  sexes  are  less  noticeable;  the  last  three  joints  of  the 
antennae  are  circular  in  section  and  stouter  in  the  male. 

Listrus  bifasciatus,  new  species. — Form  elongate,  parallel  to  ovate, 
moderately  convex.  Color  black,  second  to  the  sixth  antennal  joint 
inclusive  piceous,  second  sometimes  paler ;  tibiae  and  tarsi  more  or  less 
piceous;  surface  scarcely  feebly  purpureo-aeneous  in  lustre  and  slightly 
shining. 

Pubescence  moderately  short,  not  very  conspicuous,  brownish-black 
hairs  about  as  abundant  as  the  paler  plumbeo-cinereous  hairs  and  arranged 
in  rather  large  maculae  which  appear  as  follows :  A  large  parascutellar 
macula,  an  humeral  elongate  one  extending  arcuately  backward  and  con- 
tinuous with  the  post-basal  at  middle  of  the  elytron,  this  macula  varies  in 
size ;  median  fascia  moderately  wide  and  distinctly  zig-zag,  scarcely  attain- 
ing the  lateral  margin,  but  extending  forward  along  the  suture,  these 
extensions  frequently  separated  and  appearing  as  post-basal  sutural 
maculae ;  subapical  fascia  similar  in  form,  or  divided  at  middle  of  each 
elytron,  the  sutural  portions  tending  to  form  a  subrhomboidal  macula  on 
the  suture ;  apical  maculae  rather  large.  The  elytral  pattern  is  suggestive 
of  that  seen  in  pardalis  Casey ;  the  pale  hairs  form  relatively  narrow  areas 
between  the  dark  fasciae  and  maculae.  Pronotal  central  macula  quite 
distinctly  constricted  at  middle  and  apparently  somewhat  different  in  the 
sexes ;  lateral  semilunar  vittae  more  or  less  distinct.  Central  area  of  the 
head  usually  darkish. 

Head  slightly  broader  than  the  pronotal  apex,  moderate  in  size, 
about  as  long  as  wide ;  front  almost  feebly  convex,  although  feebly  and 
broadly  impressed  anteriorly,  punctures  rather  coarse  and  close,  slightly 
impressed,  intervals  somewhat  indented  at  the  periphery,  a  small  glabrous 
and  slightly  convex  area  at  the  epistoma.  Eyes  rather  large.  Antennae 
long,  almost  quite  similar  in  the  sexes,  very  feebly  compressed,  rather 
stout,  very  gradually  incrassate,  outer  joints  distinctly  heavier;  first  very 
stout ;  second  oval  and  a  little  longer  than  wide ;  third  obconically  sub- 
triangular,  almost  twice  as  long  as  wide ;  fourth  wider  than  the  fifth  or 
sixth,  slightly  elongate,  triangular  and  angulate  anteriorly,  and  with  the 
fifth  noticeably  compressed ;  fifth  similar  to  the  fourth  but  obtuse 


150      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

anteriorly ;  sixth  about  as  long  as  wide  and  obtusely  rounded  anteriorly ; 
seventh  and  eighth  subtriangular,  about  as  long  as  wide,  almost  circular 
in  transverse  section ;  ninth  and  tenth  similar  in  form  but  slightly  longer ; 
eleventh  less  than  twice  as  long  as  wide,  slightly  obovate,  obtusely  rounded 
at  apex.  All  joints  beyond  the  third  are  quite  abundantly  set  with  rather 
stiff  and  moderately  short  hairs. 

Pronotum  about  a  fourth  wider  than  long;  apex  arcuato-truncate, 
apical  angles  broadly  rounded  into  the  sides,  the  latter  broadly  arcuate  in 
basal  half,  thence  less  so  and  converging  moderately  to  apex,  serrules  short 
and  rather  broad,  not  very  distinct ;  base  broadly  arcuate,  rounding  into 
the  sides,  angles  absent ;  disk  widest  behind  the  middle,  quite  evenly  con- 
vex, somewhat  declivous  antero-laterally,  punctures  moderate  in  size, 
well  defined,  separated  by  a  distance  equal  to  one  or  two  times  their 
diameters,  in  central  third  the  intervals  are  flat,  glabrous,  not  noticeably 
indented,  lateral  thirds  strongly  indentato-reticulato-punctate. 

Elytra  elongate,  punctures  rather  smaller  than  those  of  the  pronotum, 
almost  arranged  in  irregular  transverse  lines  of  twos  or  fours,  separated 
by  a  distance  equal  to  one-half  to  three  times  their  diameters,  intervals 
between  the  punctate  lines  more  or  less  feebly  and  transversely  impressed, 
surface  very  finely  rugulose. 

Abdomen  very  finely  and  almost  thickly  punctate;  sixth  ventral  seg- 
ment usually  visible ;  apex  with  rather  long  and  slightly  blackish 
pubescence ;  fifth  ventral  unmodified  on  the  disk  in  the  male. 

Legs  moderate  in  length  and  stoutness ;  tarsi  long  and  slender. 

Male. — Narrow  and  parallel.  Second  and  third  antennal  joints 
stouter  than  in  the  female ;  pronotal  hourglass-shaped  macula  with  narrow 
median  lines  of  pale  hairs  at  apex  and  base ;  fifth  ventral  segment  broad 
at  apex  and  truncate. 

Female. — Elongate  ovate,  sides  rather  feebly  arcuate,  second  and 
third  antennal  joints  less  stout;  pronotal  central  macula  usually  without 
median  pale  lines ;  fifth  ventral  broadly  rounded  at  apex. 

Measurements. — Length  (Types),  3-3.6  mm.;  width,  1-1.4  mm. 

Holotype,  male,  and  paratypes  in  my  own  collection ;  allotype,  female, 
and  paratypes  in  the  collection  of  the  California  Academy  of  Sciences. 

Type  locality. — Vine  Hill,  near  Martinez,  Contra  Costa  County, 
California.  Collected  by  myself  on  November  28th,  1908.  About  twenty 
specimens  studied. 

Distribution. — California. — (Besides  at  the  type  locality,  specimens 
have  been  seen  from  Alhambra  Valley,  Contra  Costa  County ;  these  were 
collected  on  December  25th,  1913;  Niles  Canyon,  Alameda  County, 
April  19th,  1914.) 


DESCRIPTION  OF  NEW   SPECIES  151 

Apparently  related  to  Udells  Casey,  from  which  it  differs  in  the  fourth 
antennal  joint  being  wider  and  more  prominent  anteriorly  than  the  fifth, 
shorter  pubescence  and  two  distinct  elytral  fasciae;  pubescence  not  dense 
and  coarse,  the  elytral  pattern  somewhat  resembles  that  seen  in  pardalis 
Casey. 

Listrus  simplex,  new  species. — Form  parallel  to  subovate  and  moder- 
ately convex.  Color  black,  more  or  less  shining-  and  with  a  slight  virido- 
aeneous  lustre ;  antennae  rufo-testaceous  throughout,  frequently  the  distal 
joints  are  more  or  less  rufo-piceous ;  tibiae  and  tarsi  more  or  less  rufous. 

Pubescence  short,  somewhat  sparse  and  rather  less  than  coarse, 
cinereous  in  color,  with  dark  fasciae  of  brownish-black  hairs  arranged  as 
follows :  A  basal  that  may  be  narrowly  interrupted  at  the  scutellum ; 
post-basal  maculae  absent  or  small  at  middle  of  each  elytron;  submedian 
fascia  rather  wide  with  edges  more  or  less  slightly  irregular;  the  sub- 
apical  likewise  rather  wide,  entire  or  interrupted  by  a  few  pale  hairs  on 
the  suture ;  apical  maculae  small  or  obsolete.  Pronotal  maculae  apparently 
more  or  less  different  in  the  sexes ;  lateral  semilunar  vittae  more  or  less 
evident,  sometimes  very  narrowly  separated  from  the  central  macula. 

Head  about  as  long  as  wide,  moderately  finely  punctate,  punctures 
slightly  sparse,  intervals  almost  smooth  or  feebly  indentato-rugulose ; 
front  slightly  concave,  impression  broad,  with  a  small  median  almost 
glabrous  convexity  at  the  epistomal  base.  Antennae  extending  to  about 
the  pronotal  base  and  quite  similar  in  the  sexes,  rather  stout  and  some- 
what incrassate,  feebly  compressed  to  about  the  tenth  joint ;  second,  third 
and  fourth  joints  subequal  in  length ;  second  oval,  about  as  wide  as  the 
fifth;  third  subcylindrical ;  fourth  feebly  angulate  anteriorly;  fifth  sub- 
triangular,  wider  than  the  two  preceding  and  following  joints,  as  wide  as 
the  ninth,  apical  angle  prominent  anteriorly ;  sixth  and  seventh  similar  in 
length  and  form,  the  former  slightly  smaller  and  obconico-triangular ; 
eighth  oblong-obconic ;  ninth  slightly  wider  and  almost  subquadrate; 
tenth  oblong-triangular  and  slightly  transverse ;  eleventh  suboblong-oval, 
about  a  half  longer  than  wide;  joints  from  the  sixth  to  the  eleventh 
gradually  increasing  in  width ;  outer  joints  rather  abundantly  set  with 
pale  hairs. 

Pronotum  a  little  wider  than  long,  strongly  convex  from  side  to  side, 
widest  behind  the  middle ;  sides  evenly  arcuate  in  basal  half,  thence  feebly 
arcuate  and  moderately  convergent  to  apex,  serrules  small ;  base  broadly 
and  slightly  arcuate,  apparently  rather  briefly  and  obliquely  sinuate 
within  the  obtusely  rounded  basal  angles ;  apical  angles  obtusely  rounded ; 
apex  broadly  and  feebly  arcuate;  disk  with  moderately  small  punctures, 
separated  by  a  distance  equal  to  their  diameter  in  the  central  area  from 


152      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

apex  to  base,  intervals  rather  smooth,  although  very  feebly  indentato- 
punctate,  densely  reticulato-punctate  laterally. 

Elytra  about  twice  as  long  as  wide,  oblong;  sides  parallel  to  feebly 
divergent  posteriorly,  apex  broadly  rounded ;  punctures  of  the  disk  rather 
coarse  and  separated  by  a  distance  equal  to  one  or  three  times  their 
diameter,  finer  and  more  widely  spaced  toward  apex. 

Abdomen  finely  and  not  noticeably  densely  punctate,  except  on  the 
fifth  segment,  which  is  unmodified  on  the  disk  in  the  male. 

Legs  slightly  elongate,  femora  subparallel. 

Male. — Rather  elongate,  parallel  and  narrower.  Pronotal  central 
macula  more  or  less  oval.  Antennae  slightly  stouter.  Abdomen  more 
coarsely  punctate  toward  the  apex ;  fifth  ventral  segment  truncate  at  apex. 

Female. — Subparallel,  stouter  and  slightly  wider  posteriorly;  sides 
very  feebly  arcuate.  Pronotal  central  macula  more  or  less  hourglass-shaped, 
anterior  lobe  about  reaching  to  the  apical  margin,  posterior  lobe  with  a 
short  reentering  median  line  of  pale  hairs ;  lateral  semilunar  lines  rather 
indefinite.  Fifth  ventral  rounded  at  apex. 

Measurements. — Length   (Types),  2.0-3.0  mm.;  width,  .8-1.1  mm. 

Type  locality. — Marsh  Ranch,  Green  Point,  Humboldt  County, 
California. 

Holotype,  male,  and  paratypes  in  my  own  collection ;  allotype,  female, 
in  that  of  the  California  Academy  of  Sciences. 

Habitat. — California  (on  Green  Point  and  Redwood  Creeks,  Hum- 
boldt County,  in  June ;  in  May,  F.  W.  Nunenmacher ;  Del  Norte  County, 
F.  W.  Nunenmacher;  Duncan  Mills,  Sonoma  County,  June,  F.  E.  Blais- 
dell). 

Number  of  specimens  studied,  29. 

Variations. — Antennae  and  legs  may  be  piceous.  The  basal  dark 
fascia  of  the  elytra  may  be  dissolved  into  a  juxta-scutellar  macula  on  each 
side,  and  a  humeral  lunule  that  may  join  the  post-basal  macula. 

Simplex  in  color  and  maculation  resembles  annulatus,  but  is  more 
elongate  and  less  stout,  with  pronotum  narrower.  In  annulatus  the  basal 
fascia  is  usually  rather  widely  interrupted  at  the  scutellum  and  suture  and 
no  variations  have  been  observed. 

Listrus  incestus,  new  species. — Form  elongate,  oblong-oval  and 
moderately  convex.  Color  black,  appendages  nigro-piceous,  second  joint 
of  the  antennae  pale,  terminal  joints  black ;  surface  somewhat  shining, 
lustre  more  or  less  feebly  purpureo-cupreous  anteriorly. 

Pubescence  short  and  decumbent,  argenteo-cinereous  in  color  and 
arranged  in  maculae  and  fasciae  of  brownish  hairs  on  the  elytra  as  fol- 
lows :  A  broken  basal  fascia,  forming  a  small  humeral  and  parascutellar 
macula  on  each  elytron,  usually  small  but  variable  in  size;  a  post-basal 


DESCRIPTION  OF  NEW  SPECIES  153 

macula  usually  extending-  forward  to  the  humeral  forming  a  lunule;  a 
rather  wide  entire  median  fascia  with  slightly  zig-zag  edges,  sometimes 
tending  to  extend  basalward  along  the  suture;  a  similar  but  usually 
slightly  narrower  subapical  fascia,  and  an  apical  varying  in  size,  some- 
times involving  the  apex.  Pronotal  median  macula  constricted  at  middle, 
frequently  with  a  short  reentrant  and  median  line  of  pale  hairs  at  apex 
and  base ;  lateral  vittae  narrow  or  broader.  Frontal  region  of  the  head 
sometimes  darkish. 

Head  about  as  long  as  wide.  Front  feebly  and  broadly  concave, 
scarcely  convex  in  the  median  line  behind  the  epistoma,  surface  densely 
indentato-punctate.  Antennae  extending  a  little  beyond  the  pronotal  base, 
not  noticeably  slender,  feebly  compressed;  second  joint  oval,  almost  as 
wide  as  long ;  third  cylindrico-obconical,  scarcely  longer  than  the  second ; 
fourth  noticeably  subangulate  anteriorly;  fifth  wider,  distinctly  angulate, 
sides  unequal,  apical  margin  oblique ;  sixth  smaller,  as  long  as .  wide, 
obtusely  rounded  anteriorly ;  seventh  oblong-triangular  and  about  as  long 
as  wide;  eighth  similar  and  slightly  thicker;  ninth  subtriangular  and 
slightly  wider  than  long ;  tenth  scarcely  wider  than  long,  sides  arcuate ; 
eleventh  oblong-oval,  apex  obtusely  rounded,  about  a  half  longer  than 
wide ;  last  four  joints  gradually  incrassate. 

Pronotum  about  a  fifth  wider  than  long,  evenly  convex ;  apex  arcuato- 
truncate,  apical  angles  distinctly  rounded;  base  broadly  arcuate,  angles 
obtuse,  small  and  more  or  less  reflexed,  sometimes  scarcely  discernible; 
sides  strongly  and  evenly  arcuate  in  basal  half,  thence  straighter  and 
moderately  converging  to  apex,  serrules  equal,  rather  thick  and  sub- 
obtuse,  almost  equally  spaced;  fimbriae  moderately  long  and  not  dense; 
disk  densely  indentato-punctate.  Sometimes  the  sides  are  very  briefly 
sinuate  before  the  basal  angles. 

Elytra  oblong-suboval,  twice  as  long  as  wide;  base  truncate,  apex 
rather  broadly  rounded ;  punctures  rather  coarse,  separated  by  a  distance 
equal  to  once  or  twice  their  own  diameter,  finer  toward  apex;  surface 
somewhat  transversely  impresso-rugose. 

Abdomen  densely  and  finely  sculptured.  Fifth  ventral  segment 
unmodified  in  the  male. 

Legs  not  stout.  Femora  subparallel ;  surface  finely  punctato- 
rugulose.  . 

Male. — Narrower.     Fifth  ventral  truncate  at  apex. 

Female.— Somewhat  broader,  slightly  wider  posteriorly.  Antennae 
similar  to  those  of  the  male,  but  rather  less  stout.  Fifth  ventral  rather 
broadly  rounded  at  apex. 

Measurements.— Length  (Types),  2.5-3.2  mm.;  width,  1-1.2  mm. 

Holotype,  male,  and  allotype,  female,  in  the  author's  collection. 


154      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Collected  by  Mr.  J.  O.  Martin,  who  possesses  paratypes. 

Type  locality  and  habitat. — Bear  Lake,  San  Bernardino  County,  Cali- 
fornia. Collected  May  5,  1919. 

Remarks. — Incestus  resembles  both  simplex,  n.  sp.,  and  incertus 
Casey.  In  incertus  the  sexes  are  shorter  and  rather  more  robust,  the  male 
being  rather  more  oblong ;  centrally  the  pronotal  disk  is  simply  punctate, 
antennae  slenderer,  joints  five  to  ten  triangular,  tibiae  and  tarsi  pale. 

Simplex  has  the  central  area  of  the  pronotum  simply  punctate ; 
antennae  more  compact,  fourth  joint  somewhat  shorter  than  the  third  and 
distinctly  obconical,  fifth  scarcely  wider  and  feebly  angulate  anteriorly; 
elytral  dark  fasciae  rather  broader  and  straighter,  tibiae,  tarsi  and  antennae 
more  or  less  pale. 

In  difficilis  the  third  joint  of  the  antennae  is  cylindrical,  the  fifth 
longer  than  wide,  distinctly  wider,  sixth  and  following  joints  distinctly 
shorter  than  the  fourth.  Central  area  of  the  pronotal  disk  not  densely 
indentato-punctate. 

Listrus  giffardi,  new  species. — Form  oblong-oval,  subparallel,  slightly 
elongate  and  very  moderately  convex.  Color  black,  upper  surface  of  body 
with  a  more  or  less  noticeable  aeneous  lustre ;  antennae  rufous,  blackish 
distally ;  legs  rufous  throughout. 

Pubescence  moderately  short,  not  coarse,  recumbent,  moderate  in 
abundance  and  plumbeo-cinereous  in  color;  slightly  longer  on  the  under 
surface  of  the  body,  scarcely  paler  anteriorly.  Elytral  dark-brownish 
hairs  arranged  in  maculae  and  fasciae  as  follows :  An  elongate  narrow 
humeral  and  a  broader  posteriorly  elongate  parascutellar  macula  on  each 
elytron ;  a  post-basal  small  macula  at  middle  on  each  side ;  submedian  and 
subapical  transverse  fasciae  rather  wide  with  edges  irregular ;  apical 
maculae  rather  large  and  distinct.  Pronotal  pubescence  similar  to  that 
on  the  elytra,  maculae  obscure,  central  constricted  figure  and  moderately 
wide  lateral  vittae  feebly  evident ;  front  of  head  darkish  in  the  central  area. 

Head  about  as  long  as  wide,  not  large,  eyes  moderately  prominent 
and  evenly  convex ;  front  feebly  and  broadly  impressed,  punctures 
moderately  coarse,  rather  sparse,  intervals  smooth  and  obsoletely 
indentate ;  muzzle  small.  Antennae  rather  long  and  somewhat  slender, 
moderately  compressed ;  second  joint  oval,  not  stout,  not  twice  as  wide 
as  the  third,  the  latter  cylindrical,  more  than  twice  as  long  as  wide,  equal 
to  the  fourth  in  length ;  the  latter  obconical,  slightly  prominent  anteriorly 
at  apex ;  fifth  about  a  half  longer  than  wide,  not  widely  triangular,  about 
as  wide  as  the  eighth  or  ninth,  apical  margin  oblique,  moderately  promi- 
nent anteriorly ;  sixth  and  seventh  subequal  in  length  and  size,  almost  a 
little  longer  than  wide,  apical  margin  oblique;  eighth  to  the  tenth  tri- 


DESCRIPTION  OF  NEW   SPECIES  155 

angular,  slightly  longer  than  wide,  increasing  very  slightly  in  width  and 
size  in  the  order  named ;  eleventh  almost  evenly  oval,  slightly  narrowed 
apically,  about  twice  as  long  as  wide. 

Pronotum  a  little  wider  than  long,  widest  just  behind  the  middle, 
length  equal  to  breadth  of  the  head  across  the  eyes ;  apex  arcuato-truncate 
in  circular  arc ;  apical  angles  rounded ;  sides  not  strongly  arcuate  in  basal 
half,  slightly  sinuate  before  the  basal  angles,  anteriorly  .rather  straight 
and  moderately  convergent  to  apex,  serrules  small,  not  very  distinct 
although  a  few  may  be  larger  and  subacute,  fimbriae  moderate  in  length  ; 
base  broadly  arcuate,  feebly  sinuate  laterally;  basal  angles  obtuse  and 
more  or  less  distinct;  disk  rather  strongly  convex,  rather  strongly 
arcuately  declivous  antero-laterally,  sides  feebly  impressed  within  the 
basal  angles  and  laterally  anteriorly,  punctures  moderate  in  size,  rather 
sparse  in  the  central  two-fourths,  intervals  flat,  rather  smooth,  apparently 
obsoletely  indentate,  lustre  rather  dull,  in  lateral  fourths  not  strongly  nor 
very  densely  reticulato-punctate. 

Elytra  oblong-oval,  twice  as  long  as  wide  and  parallel ;  humeri  not 
prominent ;  disk  rather  distinctly  transversely  impressed  behind  the  base, 
punctures  rather  indentated  and  thereby  appearing  coarse,  separated  by 
rather  less  than  twice  their  own  width  and  slightly  arranged  in  transverse 
rows,  with  the  intervals  feebly  and  transversely  impressed  giving  the  sur- 
face the  appearance  of  being  subrugose,  surface  smoother  apically  with 
the  punctures  finer ;  apex  evenly  but  not  very  broadly  rounded. 

Abdomen  very  finely  and  densely  punctulate  and  microscopically 
reticulate,  less  than  moderately  convex. 

Legs  moderate  in  length  and  stoutness. 

Salient  male  characters. — Subparallel  and  elongate,  moderately 
narrow;  sides  of  the  fifth  ventral  segment  moderately  convergent  pos- 
teriorly, apex  equal  in  width  to  about  a  third  of  the  base  and  truncate  at 
tip ;  surface  not  in  the  least  modified,  apical  tactile  setae  noticeably  few. 

Female  unknown. 

Measurements. — Length,  3  mm. ;  width,  1  mm. 

Holotype,  male,  in  my  own  collection ;  collected  on  June  9,  1917,  by 
Mr.  Walter  M.  Giffard  of  Honolulu,  Hawaii. 

Type  locality. — Santa  Cruz  County,  at  an  elevation  of  600  feet. 

Giffardi  is  peculiar  in  its  sparsely  punctured  pronotum  and  entirely 
pale  legs,  the  elytral  maculation  is  different  from  that  observed  in  luteipes 
to  which  it  is  apparently  related,  the  fasciae  being  more  sharply  defined 
and  the  basal  maculae  sharper  and  more  elongate,  and  besides  the 
pronotum  is  narrower  than  in  that  species  where  the  male  is  more  ovate  in 
form.  I  take  pleasure  in  naming  this  species  after  its  discoverer,  Mr. 
Giffard. 


156      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Listrus  parvicollis,  new  species. — Form  elongate  subovate,  moderately 
convex.  Color  black  with  a  faint  purpureo-aeneous  lustre ;  labrum  more 
or  less  rufous,  head  and  pronotum  opaque,  antennae  piceo-rufous,  joints 
two,  three  and  four  more  rufous  than  the  terminal  ones,  elytra  more  or 
less  shining;  femora  rufo-piceous,  tibiae  and  tarsi  rufous. 

Pubescence  short  and  sparse,  dark  cinereous  in  color  with  brownish 
hairs  arranged  in  a  pattern  as  follows:  A  broad  median  fascia  that 
extends  along  the  suture  to  base  and  dilating  about  the  scutellum,  the 
humeral  and  lateral  area  with  irregular  grayish  markings ;  a  wide  sub- 
apical  fascia  that  involves  most  of  the  apex,  where  there  are  scattered 
grayish  hairs.  The  median  and  subapical  fasciae  are  separated  by  a  very 
narrow  transverse  zig-zag  fascia  of  pale  hairs.  Pronotal  central  macula 
very  faintly  indicated,  but  apparently  constricted  at  middle  and  with  a 
narrow  median  grayish  line;  lateral  semilunar  vittae  more  or  less 
obsolescent. 

Head  notably  small,  about  as  wide  as  long,  muzzle  small  and  rela- 
tively narrow ;  front  moderately  impressed,  impressions  broad  and  sepa- 
rated by  a  feeble  but  distinct,  glabrous  raised  area  behind  the  base  of  the 
epistoma;  surface  densely  punctato-rugose,  eyes  rather  prominent. 
Antennae  extending  to  the  base  of  the  pronotum,  rather  slender  and  some- 
what incrassate ;  second  joint  oval  and  rather  stout ;  third  cylindric  and 
about  twice  as  long  as  wide ;  fourth,  fifth  and  sixth  distinctly  angulate  on 
their  anterior  borders ;  fifth  distinctly  wider  than  the  fourth  but  subequal 
in  width  with  the  sixth;  seventh,  eighth,  ninth  and  tenth  not  stout,  but 
evidently  slightly  transverse ;  eleventh  oval  and  obtusely  pointed  and 
about  a  half  longer  than  wide. 

Pronotum  notably  small,  about  a  third  wider  than  long ;  sides  evenly 
and  rather  strongly  rounded  in  basal  half,  thence  straight  and  convergent 
to  apex,  and  coinciding  with  the  inner  border  of  the  eyes,  serrules  small 
and  acute,  fimbriae  not  close  and  moderately  short ;  apical  angles  obtuse ; 
apex  rather  arcuate  and  distinctly  narrower  than  the  head ;  base  broadly 
rounded,  continuously  so  with  the  sides ;  disk  very  densely  indentato- 
punctate,  indentations  same  size  as  the  punctures,  the  deeper  punctures 
separated  by  one  or  two  indentations ;  surface  rather  strongly  and  evenly 
convex  from  side  to  side,  especially  anteriorly. 

Elytra  distinctly  wider  than  the  pronotum,  slightly  more  than  twice 
as  long  as  wide;  sides  broadly  and  not  strongly  arcuate;  apex  rather 
broadly  rounded ;  disk  not  very  coarsely  punctate,  punctures  separated  by 
a  distance  about  equal  to  twice  their  own  diameter,  finer  toward  apex. 

Abdomen  very  finely  punctate  and  more  or  less  finely  reticulate. 

Male  unknown. 


DESCRIPTION  OF  NEW  SPECIES  157 

Female. — Fifth  ventral  abdominal  segment  rounded  at  apex,  tip 
deflexed,  surface  with  a  small  but  distinct  impression  in  front  of  the 
deflexed  margin. 

Measurements. — Length,  2.4  mm. ;  width,  .9  mm. 

Holotype,  female,  in  my  own  collection. 

Type  locality. — Mokelumne  Hill,  Calaveras  County.  Collected  in 
May.  A  single  specimen  studied.  A  very  distinct  species  in  the  charac- 
ters of  the  head,  pronotum  and  elytral  maculation. 

Listrus  trochantcricus,  new  species. — Form  slightly  robust,  oblong- 
ovate,  rather  more  than  twice  as  long  as  wide.  Color  black;  tibiae  and 
tarsi  rufo-testaceous,  tibiae  at  base  and  tarsi  toward  tip,  blackish ;  second 
joint  of  the  antennae  rufo-testaceous,  first  joint  black  as  usual,  remaining 
joints  rufo-piceous,  last  two  or  three  may  be  quite  black. 

Pubescence  rather  dense  and  coarse,  almost  concealing  the  general 
surface,  cinereous  in  color,  with  brownish-black  maculae  and  fasciae 
arranged  on  each  elytron  as  follows:  A  rounded  parascutellar  macula 
and  a  humeral  lunule  more  or  less  interrupted  behind  the  umbo, 
umbonal  dot  small  or  absent,  apical  portion  feebly  arcuate,  appearing  as  a 
post-basal  macula ;  a  narrow  transverse  fascia  a  little  behind  the  middle, 
slightly  zig-zag,  and  interrupted  at  the  suture,  often  broken  into  two 
maculae  on  each  elytron ;  subapical  transverse  fascia  similar  in  form ; 
apical  macula  present  or  absent.  Pronotal  pattern  consists  of  lateral 
semilunar  vittae  and  a  somewhat  constricted  central  figure,  the  former 
broken  into  two  and  the  latter  more  or  less  dissolved  into  four  parts. 
Central  area  of  the  head  submaculate.  Maculae  of  the  head  and  pronotum 
not  very  distinct. 

Head  slightly  transverse,  front  rather  broadly  impressed,  or  longi- 
tudinally impressed  within  the  eyes,  with  a  feeble  central  longitudinal  con- 
vexity, punctures  not  coarse,  rather  crowded  in  the  impressions,  some- 
what sparse  elsewhere,  intervals  scarcely  indented,  a  small  impunctate  area 
at  base  of  the  epistoma ;  muzzle  short,  margin  evenly  arcuate  from  side  to 
side.  Antennae  rather  slender,  feebly  incrassate,  outer  joints  noticeably 
heavier,  slightly  compressed ;  third  cylindrical,  fourth  obconico-triangular, 
both  slightly  elongate ;  fifth  serrate,  a  little  wider  than  the  sixth,  sixth  to 
the  tenth  inclusive,  subtriangular. 

Pronotum  less  than  a  fourth  wider  than  long ;  sides  evenly  and  rather 
strongly  arcuate,  moderately  convergent  anteriorly  and  less  arcuate, 
serrules  not  strong ;  basal  angles  not  evident  and  well  rounded ;  base 
broadly  arcuate ;  disk  evenly  convex,  more  strongly  so  anteriorly,  closely 
punctate,  punctures  moderate  in  size,  separated  by  a  distance  more  or  less 
equal  to  their  diameter,  intervals  scarcely  indented  in  the  central  area,  but 


158      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

densely  indentato-punctate  in  the  lateral  two-fifths,  lustre  dull ;  fimbriae 
moderate  in  length,  even  and  cinereous  in  color. 

Elytra  oblong,  less  than  twice  as  long  as  wide  and  a  little  wider 
posteriorly ;  sides  feebly  arcuate,  subparallel,  moderately  convex  on  the 
disk;  punctures  separated  by  a  distance  equal  to  about  twice  their 
diameter,  at  base  subequal  to  those  of  the  pronotum,  finer  apically  and 
more  widely  spaced ;  apex  rather  broadly  rounded. 

Abdomen  finely  and  rather  sparsely  punctate;  surface  extremely 
finely  rugulose ;  pubescence  slightly  longer  and  finer.  Fifth  ventral  seg- 
ment modified  on  the  disk  in  the  male. 

Legs  moderate  in  length  and  stoutness.  Metatrochanters  triangular ; 
femora  quite  cylindrical  behind  the  trochanters  and  more  or  less  arcuate. 

Male. — Narrower.  Metatrochanters  subacute  on  their  inner  angle ; 
femora  more  strongly  cylindrical  and  arcuate  behind  the  trochanters. 
Fifth  ventral  segment  shorter  and  evidently  sinuato-truncate  at  apex; 
hairs  of  the  genital  segment  more  or  less  brownish  or  blackish. 

Female. — Broader.  Metatrochanters  rounded  on  their  inner  angle ; 
femora  less  cylindrical  and  arcuate  at  base ;  fifth  ventral  longer  with  the 
margin  broadly  rounded  at  apex. 

Measurements. — Length  (Types),  2.5-3  mm.;  width,  .9-1.1  mm. 

Holotype,  male,  and  allotype,  female,  in  my  own  collection.  Para- 
types  in  my  own  and  that  of  Mr.  J.  O.  Martin. 

Type  locality. — Bear  Lake,  San  Bernardino  County,  California. 
Collected  by  Mr.  J.  O.  Martin,  on  May  5,  1919.  A  series  of  twenty 
specimens  studied. 

Tro chant ericus  differs  from  all  known  species  in  the  form  of  the 
metatrochanters  and  metafemora.  The  elytral  fasciae  are  usually  broken 
up  into  rounded  or  slightly  transverse  maculae,  never  attaining  the  suture 
in  the  series  at  hand.  The  central  dark  figure  of  the  pronotum  may  be 
broken  up  or  shaped  like  two  U's  with  their  bases  together. 

Listrus  liebecki,  new  species. — Form  distinctly  oblong-oval  and 
moderately  convex.  Color  very  black,  with  a  more  or  less  dark  virido- 
metallic  lustre;  basal  joints  of  the  palpi  and  mandibles  more  or  less  rufo- 
piceous ;  second  antennal  joint  rufous,  the  four  following  joints  more  or 
less  dark  rufo-piceous ;  tarsi  and  distal  half  of  the  tibiae  more  or  less  dark 
rufous  to  rufo-piceous. 

Pubescence  sparse,  short,  closely  recumbent  and  very  inconspicuous, 
dark  plumbeous  in  color,  dark  areas  clothed  with  very  dark-brown  hairs. 
Maculation  very  obscure  and  apparently  consists  of  rather  broad  fasciae 
and  maculae  arranged  as  follows :  A  large  parascutellar  and  a  smaller 
humeral  macula  on  each  elytron ;  a  post  basal  macula  at  middle  of  each ; 


DESCRIPTION  OF' NEW   SPECIES  159 

a  median  and  subapical  fascia,  the  latter  very  indefinite ;  apical  maculae 
not  discernible  in  the  type.  Pronotal  central  macula  apparently  oval, 
lateral  semilunar  vittae  not  discernible.  Hairs  on  the  head  more  or  less 
diffusely  brownish ;  those  on  the  prosternum  longer  and  more  cinereous, 
elsewhere  beneath  similar  in  length  but  somewhat  darker. 

Head  slightly  transverse,  distinctly  wider  than  the  pronotal  apex ; 
eyes  moderately  large  and  prominent;  front  broadly  impressed,  a  small 
and  convex,  median  and  more  or  less  glabrous  area  just  behind  the 
epistomal  base;  punctures  rather  coarse,  well  defined  and  rounded, 
intervals  more  or  less  indented.  Antennae  moderately  long,  slightly 
stout,  scarcely  incrassate;  second  joint  oval,  about  a  half  longer  than 
wide,  rather  less  than  half  the  width  of  the  first;  third  joint  cylindrical, 
scarcely  longer  than  the  second,  slightly  widest  at  apex ;  fourth  feebly 
compressed,  subtriangular,  not  prominent  anteriorly,  about  a  half  longer 
than  wide ;  fifth  slightly  compressed,  subtriangular,  about  a  fourth  longer 
than  wide,  wider  than  the  four  following  joints,  apical  angle  prominent 
anteriorly;  sixth,  seventh,  and  eighth  subtriangular,  less  compressed, 
about  as  long  as  wide ;  ninth  similar  in  form  and  a  little  longer ;  tenth 
slightly  stouter,  more  oblong,  very  slightly  longer  than  wide;  eleventh 
almost  oblong-oval,  rather  less  than  twice  as  long  as  wide,  not  strongly 
narrowed  at  apex. 

Pronotum  about  a  fourth  wider  than  long,  rather  evenly  convex ;  apex 
arcuato-truncate ;  apical  angles  rather  broadly  rounded  into  the  sides,  the 
latter  broadly  arcuate,  rather  more  strongly  so  in  basal  half,  converging 
moderately  to  apex,  serrules  small,  rather  widely  spaced,  subacute, 
fimbriae  moderate  in  length  and  plumbeo-cinereous  in  color ;  base  broadly 
arcuate  and  not  strongly  lobed,  sinuate  laterally ;  basal  angles  obtuse  and 
distinct;  disk  moderately  coarsely  punctate  in  the  central  area,  intervals 
about  equal  to  the  diameter  of  the  punctures,  indentato-punctate  apically 
and  basally,  laterally  very  densely  indentato-reticulato-punctate. 

Elytra  oblong,  about  twice  as  long  as  wide,  base  rather  broadly 
emarginate,  scutellum  transversely  oblong ;  disk  rather  coarsely  punctate, 
punctures  somewhat  shallow,  separated  by  a  distance  equal  to  one  or  two 
times  their  diameter,  a  few  coalescing  transversely  producing  a  somewhat 
but  slight  rugoseness,  punctures  finer  toward  apex,  the  latter  rather  para- 
bolically  rounded. 

Abdomen  rather  densely  punctate ;  fifth  ventral  segment  modified  in 
the  male. 

Legs  rather  short  and  moderate  in  stoutness. 

Male. — Fifth  ventral  distinctly  modified,  apex  rather  deeply  sinuate, 
adjacent  surface  somewhat  impressed,  impression  limited  by  denser  punc- 
tuation, hairs  slightly  coarser,  but  pale  in  color,  or  at  least  very  slightly 


160      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

darker  about  the  abdominal  apex;  metafemora  thickened  and  apparently 
arcuate,  sides  at  apical  third  moderately  swollen. 

Female  unknown. 

Measurements. — Length,  3.2  mm.;  width,  1.1  mm. 

Holotype,  male,  in  my  own  collection ;  paratypes  in  that  of  Mr.  Chas. 
Liebeck. 

Type  locality. — Fort  Collins,  Colorado.  Only  known  from  the  type 
locality. 

Remarkably  distinct  from  interruptus  and  the  unicolorous  species  of 
the  same  region.  It  has  not  been  possible  to  study  the  metatrochanters  on 
account  of  their  being  hidden  by  glue  and  the  lack  of  specimens  for 
dissection. 

Listrus  olympianus,  new  species. — Form  elongate  suboval,  slightly 
robust  and  rather  strongly  convex.  Color  deep  black  with  a  slight  bluish 
metallic  lustre ;  head  and  pronotum  feebly  purpureo-aeneous ;  tarsi  and 
tips  of  the  tibiae  somewhat  rufo-piceous ;  second  antennal  joint  more  or 
less  rufous,  third  joint  rufo-piceous. 

Pubescence  of  less  than  moderate  length,  rather  sparse,  mouse- 
colored  and  in  certain  lights  flavo-plumbeous,  dark  areas  difficult  of 
determination.  Basal  fascia  broken  into  two  maculae  on  each  elytron, 
the  humeral  small  and  the  parascutellar  larger ;  post-basal  represented  by 
an  indefinite  macula  at  middle  of  each  elytron ;  median  fascia  irregularly 
and  obscurely  transverse ;  subapical  fascia  transverse,  entire  or  broken 
and  rather  indefinite ;  apical  maculae  more  or  less  obscure.  Maculations 
are  moderately  wide  as  a  rule.  Pronotal  median  macula  constricted  at 
middle,  faintly  incised  anteriorly  and  posteriorly  on  the  median  line; 
lateral  semilunar  vittae  distinct  or  more  or  less  atrophic.  Pronotal 
fimbriae  not  long  nor  close  and  cinereous  in  color. 

Head  rather  large  and  slightly  wider  than  the  pronotal  apex ;  front 
broadly  impressed  between  the  eyes,  a  feeble  median  longitudinal  con- 
vexity is  discernible;  rather  closely  and  distinctly  punctate,  intervals 
obsoletely  indentate,  antennal  convexities  punctato-rugose ;  muzzle  short 
with  sides  convergent.  Antennae  reaching  to  about  the  pronotal  base, 
not  distinctly  incrassate  and  dissimilar  in  the  sexes,  slightly  compressed, 
clothed  rather  thickly  with  short,  stiff  grayish  hairs ;  joints  rather  densely 
punctate. 

Pronotum  about  a  fourth  wider  than  long,  very  slightly  narrower 
than  the  elytral  base ;  sides  rather  evenly  and  moderately  arcuate  in  basal 
half,  less  so  anteriorly  and  slightly  converging  and  rounding  into  the  apex, 
the  latter  arcuato-truncate  at  middle,  serrules  rather  coarse ;  basal  angles 
obtusely  rounded;  base  arcuate;  disk  moderately  strongly  convex,  less  so 


DESCRIPTION  OF  NEW  SPECIES  161 

across  the  base,  more  declivous  antero-laterally,  punctures  rather  coarse, 
separated  by  a  distance  about  equal  to  their  diameter,  intervals  smooth 
and  shining,  in  the  central  area  obsoletely  indentate,  not  widely  indentato- 
punctate  along  apex  and  base,  laterally  rather  widely  and  densely 
indentato-rugoso-punctate ;  apex  slightly  narrower  than  the  base. 

Elytra  about  twice  as  long  as  wide ;  sides  parallel,  very  feebly  arcuate, 
somewhat  wider  posteriorly ;  apex  rather  broadly  rounded,  sutural  angles 
obtusely  and  narrowly  rounded ;  punctures  rather  coarse,  separated  by  a 
distance  two  or  three  times  their  diameter,  surface  very  finely  crinkled. 

Abdomen  finely  punctate ;  fifth  segment  quite  densely  so  and  different 
in  the  sexes. 

Legs  moderate  in  length  and  stoutness. 

Male. — Similar  in  form  to  the  female  although  slightly  narrower. 
Antennae  stouter,  second  joint  globular;  third  narrowest  and  obconical, 
about  equal  in  length  to  the  fourth ;  the  latter  triangular,  distinctly  wider, 
equal  in  width  to  the  sixth  or  seventh,  apex  transverse ;  fifth  slightly  longer 
and  subtriangular,  broader  than  the  fourth,  sixth  or  seventh,  anteriorly 
prominent;  sixth  and  seventh  about  as  long  as  wide,  subtriangulo- 
obconical ;  eighth  rather  more  triangular  and  a  little  longer ;  ninth  about 
as  long  as  wide,  rather  subtriangular  and  with  sides  more  arcuate;  tenth 
almost  as  wide  as  long,  widest  joint  of  all  but  not  suddenly  so,  almost 
quadrate-triangular  in  outline;  eleventh  narrower,  about  as  wide  as  the 
ninth,  a  half  longer  than  wide,  tapering  gradually  in  apical  half  and  almost 
subfusiform.  Fifth  ventral  segment  moderately  broad  at  apex,  distinctly 
sinuato-truncate,  adjacent  surface  rather  glabrous  and  impressed,  sides  of 
the  impression  rather  more  prominent,  pubescence  longer,  somewhat 
denser  and  more  conspicuous,  in  part  blackish  at  margins  and  on  the 
genital  segment ;  longer  tactile  hairs  rather  abundant. 

Female. — Scarcely  stouter.  Antennae  moderately  slender,  second 
joint  slightly  less  globular;  third  subtriangular  and  comparatively  narrow; 
fourth,  sixth,  seventh  and  eighth  subtriangular,  slightly  longer  than  wide ; 
fifth  slightly  longer  than  wide,  subtriangular,  apex  rather  oblique  and 
distinctly  angulate  anteriorly,  scarcely  as  wide  as  the  ninth;  ninth  and 
tenth  subtriangular,  about  as  long  as  wide,  sides  of  the  tenth  moderately 
arcuate ;  eleventh  obovate,  wider  than  the  tenth,  about  a  half  longer  than 
wide.  Fifth  ventral  rather  broadly  rounded  at  apex.  Pubescence  longer, 
denser  and  darker  about  the  abdominal  apex. 

Measurements. — Length  (Types),  3.1-3.4  mm.;  width,  1.0-1.2  mm. 

Holotype,  male,  and  allotype,  female,  in  my  own  collection ;  paratypes 
in  that  of  Mr.  Chas.  Liebeck. 

Type  locality. — Olympia,  Washington  State.  Eight  specimens 
studied. 


162      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Remarks. — Olympianus  is  evidently  rather  closely  related  to  dilutus, 
from  which  it  differs  in  the  stouter  and  shorter  antennae,  flatter  and 
scarcely  impressed  front,  and  in  the  male  the  fifth  ventral  is  modified. 
In  dilutus  the  same  segment  is  squarely  truncate  and  narrowly  beveled  on 
the  margin  in  middle  third.  The  maculae  in  dilutus  tend  to  transverse 
fasciae,  in  both  they  are  equally  difficult  of  determination.  In  liebecki 
the  second  antennal  joint  is  elongate,  in  olympianus  it  is  globular — as  long 
as  wide. 

Listrus  occidens,  new  species. — Form  comparatively  small,  oblong- 
oval,  subparallel  and  moderately  convex.  Color  black,  with  a  distinct 
aeneous  lustre ;  mouth  parts  more  or  less  rufo-testaceous,  palpi  more  or 
less  black ;  antennae,  tibiae  and  tarsi  rufo-testaceous ;  femora  at  times 
slightly  rufescent  apically. 

Pubescence  moderately  short,  abundant  but  not  completely  hiding  the 
body  surface,  recumbent,  plumbeo-cinereous  in  color ;  longer  on  the  under 
surface  of  the  body,  snowy  white  on  the  head,  sterna  and  their  side  pieces  ; 
slightly  darker  on  the  abdomen  but  snowy  on  the  apex.  Dark  brown 
hairs  of  the  upper  surface  reduced  to  a  minimum,  maculation  of  the  elytra 
nearly  obsolete;  a  basal  parascutellar  dot  and  a  very  small  post-basal 
macula  at  middle  of  each  elytron ;  submedian  and  subapical  fasciae  repre- 
sented by  indefinite  small  maculae,  apical  dots  obsolete.  Pronotal  maculae 
indefinite,  central  figure  apparently  constricted  at  middle ;  lateral  vittae 
broken  into  two  dots  each,  the  anterior  of  which  appears  as  a  rounded 
macula  in  the  antero-lateral  quadrant  of  the  disk. 

Head  about  as  long  as  wide,  eyes  large  and  rather  strongly  convex ; 
front  broadly  impressed,  feebly  convex  on  the  vertex,  punctures  moder- 
ately large,  rather  sparse,  denser  toward  the  epistoma,  intervals  feebly 
rugulose  and  rather  shining;  white  hairs  on  the  muzzle  and  anterior 
canthi  of  the  eyes  rather  conspicuous.  Antennae  moderately  long, 
extending  to  about  the  pronotal  base,  three  apical  joints  rather  stout  and 
quite  circular  in  transverse  section,  intermediate  joints  moderately  com- 
pressed; second  joint  oval,  about  as  long  as  wide;  third  slender  and 
cylindrical,  about  as  long  as  the  second  or  the  fourth,  the  latter  sub- 
triangular  and  rather  prominent  anteriorly;  fifth  slightly  elongate, 
apparently  constricted  at  base  and  as  wide  as  the  ninth,  anterior  margin 
prominent  and  arcuately  rounded;  sixth  and  seventh  triangular,  sub- 
equal  and  as  long  as  wide;  eighth  apparently  wider  than  long  and  sub- 
triangular;  ninth  subtriang.ular  with  sides  somewhat  arcuate;  ninth  and 
tenth  a  little  stouter,  about  as  long  as  wide,  sides  arcuate ;  eleventh  short 
ovate,  less  than  a  half  longer  than  wide. 

Pronotum  about  a  fourth  wider  than  long,  widest  behind  the  middle ; 
apex  feebly  arcuate  in  circular  arc ;  sides  evenly  and  rather  moderately 


DESCRIPTION  OF  NEW  SPECIES  163 

strongly  arcuate  in  posterior  half,  thence  rather  straight  and  somewhat 
strongly  convergent  to  apex  as  viewed  from  above,  serrules  small  and 
subacute,  fimbriae  cinereous  and  moderate  in  length;  apical  angles 
obtusely  rounded;  base  broadly  arcuate,  feebly  sinuate  laterally;  basal 
angles  represented  by  a  denticle,  otherwise  rounded;  disk  evenly  and 
moderately  convex  posteriorly,  arcuately  and  strongly  declivous  antero- 
laterally ;  punctures  in  the  central  area  moderate  in  size  and  separated  by 
a  distance  equal  to  about  one  or  two  times  their  diameter,  intervals  slightly 
rugulose,  or  very  obsoletely  indentate,  laterally  not  very  coarsely 
reticulato-punctate. 

Elytra  oblong,  about  twice  as  long  as  wide,  rather  evenly  convex 
from  side  to  side ;  disk  moderately  closely  punctate,  punctures  not  strongly 
outlined,  separated  by  a  distance  equal  to  one  or  two  times  their  diameter, 
surface  somewhat  feebly  transversely  rugose;  punctures  finer  and  more 
widely  spaced  toward  apex,  the  latter  evenly  rounded. 

Abdomen  finely  and  densely  punctured ;  fifth  ventral  segment  modified 
on  the  disk  in  the  male. 

Legs  moderate  in  length  and  stoutness ;  metafemora  somewhat  con- 
stricted at  base. 

Male. — Parallel,  slightly  elongate.  Fifth  ventral  distinctly  sinuate, 
slightly  blackened  on  the  impressed  area,  hairs  pale  and  normal  elsewhere. 

Female  unknown. 

Measurements. — Length,  2.6  mm. ;  width,  .9  mm. 

Holotype,  male,  in  my  own  collection. 

Type  locality. — San  Diego,  California,  without  other  data.  Collected 
by  Mr.  O.  N.  Sanford. 

Occident  is  quite  distinct  in  its  feebly  marked  elytral  maculation, 
rufous  antennae,  tibiae  and  tarsi,  modified  fifth  ventral  segment  of  the 
abdomen,  ashy  pubescence  and  several  other  minor  differences. 

Listrus  angulatus,  new  species. — Form  oblong-ovate,  about  three 
times  as  long  as  wide  and  moderately  convex.  Color  black,  feebly 
shining,  with  a  very  slight  aeneous  lustre  anteriorly ;  tarsi  and  distal  part 
of  the  tibiae  dark  rufo-piceous ;  antennae  nigro-piceous  throughout. 

Pubescence  abundant,  conspicuous,  moderate  in  length  and  coarse- 
ness, recumbent,  longer  on  the  under  surface  of  the  body,  plumbeo- 
cinereous  in  color  and  somewhat  silvery  on  oblique  inspection.  The  dark 
brownish  hairs  are  arranged  in  more  or  less  discrete  maculae  as  follows : 
Basal  maculae  obsolete,  or  a  small  humeral  and  a  parascutellar  which  in 
some  instances  coalesce  on  each  elytron  remaining  rather  widely  inter- 
rupted at  the  scutellum ;  post-basal  macula  variable  in  size  at  middle  of 
each  elytron ;  a  submedian  fascia  which  is  very  narrow  and  zig-zag,  or 


164      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

broken  into  two  maculae  on  each  side,  of  which  the  lateral  may  be 
obsolete;  a  subapical  transverse  row  of  four  maculae  that  seldom  unite 
on  the  suture ;  apical  maculae  obsolete  or  variable  in  size.  Pronotal 
central  figure  dissolved  into  four  maculae  that  may  unite  more  or  less  to 
form  lobes  or  an  almost  constricted  macula;  lateral  vitta  broken  into  two 
dots,  it  is  seldom  entire. 

Head  relatively  small,  wider  than  the  pronotal  apex,  about  as  long  as 
wide;  eyes  large,  only  moderately  prominent;  front  broadly  and  not 
strongly  impressed,  feebly  convex  on  the  vertex,  punctures  scarcely 
coarse,  dense  anteriorly  and  more  widely  spaced  on  the  vertex,  muzzle 
short.  Antennae  similar  in  the  sexes,  moderate  in  length,  feebly  com- 
pressed, apical  joints  somewhat  circular  in  transverse  section,  joints  four 
and  five  noticeably  compressed;  second  joint  rather  stout,  subglobular, 
very  slightly  longer  than  wide,  about  as  wide  as  the  eighth;  third  about 
as  long  as  the  second,  about  half  as  wide,  obconico-cylindrical ;  fourth 
subtriangular,  slightly  longer  than  wide,  apical  margin  oblique,  prominent 
anteriorly ;  fifth  nearly  as  wide  as  the  tenth,  subtriangular,  slightly  longer 
than  wide,  apical  margin  slightly  oblique,  anterior  margin  prominent  but 
not  sharply  angulate ;  sixth  to  the  ninth  similar  in  form,  subtriangulo- 
moniliform  and  subequal  in  length ;  tenth  stouter,  about  as  long  as  wide, 
subtriangular  in  outline  as  viewed  from  above ;  eleventh  scarcely  twice  as 
long  as  wide,  obovate,  equal  in  width  to  the  tenth. 

Pronotum  about  a  third  wider  than  long,  rather  evenly  convex,  a  little 
more  declivous  antero-laterally ;  apex  subtruncate  in  moderately  circular 
arc;  sides  broadly  arcuate,  less  so  anteriorly  and  moderately  convergent 
to  apex,  serrules  small,  subacute  and  rather  widely  spaced,  fimbriae 
moderate  in  length  and  coarseness,  not  very  close;  apical  angles  sub- 
obtusely  rounded;  base  broadly  arcuate,  almost  moderately  sinuate 
laterally  within  the  angles,  which  are  evident  but  not  strong ;  disk  dull  in 
lustre,  at  center  not  very  densely  asperately  indentato-punctate,  strongly 
so  laterally. 

Elytra  oblong-oval,  humeri  rather  prominent ;  sides  rather  moderately 
arcuate,  apex  rather  broadly  rounded ;  disk  not  strongly  punctate,  punc- 
tures rather  shallow,  separated  by  a  distance  equal  to  one  or  two  times 
their  diameter,  finer  and  sparser  toward  apex. 

Abdomen  densely  and  finely  punctate,  surface  rather  microreticulate ; 
fifth  ventral  segment  modified  on  the  disk  in  the  male. 

Legs  somewhat  slender,  femora  rather  moderate  in  stoutness. 

Male. — Subparallel,  oblong-subovate ;  fifth  ventral  segment  rather 
short  and  broad,  distinctly  sinuate  at  apex,  lateral  angles  moderately 
rounded,  sinus  rather  feebly  rounded  at  bottom  where  the  edge  is  slightly 
beveled ;  adjacent  surface  more  or  less  glabrous  and  the  surrounding 


DESCRIPTION  OF  NEW  SPECIES  165 

hairs  not  modified,  although  brownish  in  color  and  slightly  denser  than 
on  the  preceding  segments. 

Female. — More  robust,  oblong-ovate;  fifth  ventral  segment  rather 
long,  only  moderately  broad  and  rather  strongly  rounded  at  apex. 

Measurements. — Length  (Types),  3.5-4.0  mm.;  width,  1.3-1.4  mm. 

Holotype,  male,  and  paratypes  in  my  own  collection ;  allotype,  female, 
in  that  of  the  California  Academy  of  Sciences. 

Type  locality. — Mono  Lake,  Mono  County,  California ;  collected  by 
Mr.  Chas.  L.  Fox,  on  June  17,  1917.  Seven  specimens  studied. 

Angulatus  differs  from  the  other  species  in  its  dark  antennae,  distinct 
basal  angles  of  the  pronotum,  stouter  ovate  form  and  simple  modification 
of  the  disk  of  the  fifth  ventral  in  the  male.  It  resembles  montanus  Casey 
in  appearance. 

Listrus  salicis,  new  species. — Form  oblong-oval,  similar  in  the  sexes 
and  moderately  convex.  Color  black,  dull  in  lustre  with  a  feeble  aeneous 
tinge;  mouth-parts  piceous;  second  joint  of  the  antennae  rufous,  third, 
fourth  and  fifth  joints  more  or  less  rufo-piceous,  apical  joints  black; 
tibiae  and  tarsi  piceous,  the  latter  may  be  rufo-piceous. 

Pubescence  rather  coarse,  linearly  subsquamiform,  rather  short  and 
recumbent,  quite  dense,  plumbeo-cinereous  on  the  upper  surface;  longer 
on  the  under  parts,  dense  and  snowy-white  on  the  prosternum  and 
sternal  side-pieces,  slightly  darker  on  the  abdomen.  The  dark  brownish 
hairs  form  rather  moderate-sized  maculae  which  are  usually  rather  obso- 
lete toward  base  but  otherwise  arranged  as  follows :  An  occasional  small 
humeral  and  parascutellar  macula;  post-basal  small  at  middle  of  each 
elytron,  or  obsolete ;  submedian  fascia  obsolete  or  very  narrow  and  zig- 
zag, or  broken  into  two  short  oblique  dots  on  elytron ;  subapical  trans- 
verse row  of  four  maculae  which  are  usually  subequal  and  discrete; 
apical  maculae  small  or  obsolete.  Pronotal  maculae  obscure,  discernible 
as  faint  clouds,  the  central  one  at  times  appears  somewhat  oblong. 

Head  about  as  long  as  wide,  eyes  large  and  moderately  strongly 
convex ;  front  broadly  and  rather  deeply  impressed  between  the  antennae 
and  anterior  canthi  of  the  eyes,  feebly  convex  toward  vertex;  surface 
rather  finely  and  densely  indentato-punctate,  punctures  coarser 'on  the 
vertex  with  intervals  rugulose;  muzzle  short  and  relatively  small;  hairs 
at  sides  of  the  front  and  before  the  eyes  rather  dense  and  white. 
Antennae  similar  in  the  sexes,  moderately  long  and  attaining  the 
pronotal  base,  rather  slender  and  moderately  compressed,  apical  three 
joints  slightly  thickened;  second  joint  oval,  slightly  longer  than  wide; 
third  feebly  obconical — nearly  cylindrical,  subequal  in  length  to  the 
fourth,  the  latter  obconico-triangular,  apical  margin  slightly  oblique, 


166      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

feebly  prominent  anteriorly  at  the  apical  angle ;  fifth  as  wide  as  the  tenth, 
triangular,  slightly  elongate,  prominent  anteriorly,  apical  margin  feebly 
oblique ;  sixth  to  the  eighth  subequal  in  width,  sixth  and  eighth  about 
equal  in  length,  sixth  subtriangular  and  as  long  as  wide ;  seventh  a  little 
longer  than  wide,  subtriangular;  eighth  about  as  long  as  wide,  less  tri- 
angular ;  ninth  about  as  wide  as  long,  sides  feebly  arcuate ;  tenth  a  little 
broader,  about  as  long  as  wide,  sides  more  arcuate,  with  the  ninth  and 
eleventh  more  circular  in  transverse  section ;  eleventh  about  a  half  longer 
than  wide,  obovate ;  pubescence  short  and  very  inconspicuous.  The 
seventh  joint  tends  to  be  slightly  wider  than  the  sixth  or  eighth. 

Pronotum  about  a  third  wider  than  long,  quite  evenly  convex,  slightly 
more  declivous  antero-laterally ;  apex  feebly  arcuate  in  circular  arc ;  sides 
broadly  arcuate,  more  strongly  so  behind  the  middle,  less  so  and  moder- 
ately convergent  anteriorly  to  apex,  serrules  small  and  inconspicuous ; 
fimbriae  moderately  short,  cinereous  and  rather  close;  apical  angles 
obtusely  rounded ;  base  broadly  arcuate  and  rather  feebly  sinuate  within 
the  very  obtuse  basal  angles,  which  are  as  a  rule  rather,  broadly  rounded ; 
disk  widest  just  behind  the  middle,  rather  coarsely  and  densely  indentato- 
punctate. 

Elytra  oblong,  twice  as  long  as  wide,  sides  parallel ;  disk  rather 
closely  and  regularly  punctate,  punctures  separated  by  a  distance  equal  to 
their  diameter,  finer  and  sparser  apically ;  surface  very  finely  micro- 
scopically rugulose ;  apex  parabolically  rounded. 

Abdomen  densely  and  finely  punctate,  pubescence  abundant  and  hiding 
the  body  surface;  fifth  ventral  segment  modified  on  the  disk  in  the  male. 

Legs  moderate  in  length ;  metafemora  not  swollen,  subparallel. 

Male. — Slightly  more  parallel.  Fifth  ventral  segment  distinctly 
sinuate,  bottom  of  the  sinus  transverse,  angles  obtuse  and  hidden  by  the 
pubescence;  adjacent  surface  impressed  and  somewhat  shining,  sub- 
glabrous,  hairs  of  the  impressed  area  and  terminal  segment  brownish 
black,  longer  and  not  noticeably  bristling,  pubescence  beyond  impressed 
area  normal.  Abdomen  but  very  slightly  longer  than  a  femur. 

Female. — Slightly  stouter,  sides  of  the  body  rather  more  arcuate. 
Abdomen  about  twice  as  long  as  a  femur ;  fifth  ventral  rather  strongly 
rounded  at  apex  and  less  broadly  so  than  in  some  of  the  other  species. 

Measurements. — Length  (Types),  3.5-4.0  mm.;  width,  1.2-1.4  mm. 

Holotype,  male,  in  my  own  collection ;  allotype,  female,  in  the  collec- 
tion of  the  California  Academy  of  Sciences ;  paratypes  in  both  collections. 

Type  locality. — Hullville,  Gravelly  Valley,  Lake  County,  California. 
Collected  by  myself  on  June  18,  1917.  Found  abundantly  on  the  catkins 
of  the  willow. 


DESCRIPTION  OF  NEW  SPECIES  167 

Solids  appears  distinct  by  its  dull  lustre,  sparse  maculation,  even  and 
rather  dense  pubescence  and  even,  close  elytral  punctuation.  Closely 
related  to  niveicanthus  which  has  a  more  pronounced  maculation.  Some- 
times salicis  is  almost  unicolorous  from  diminution  of  the  maculae. 

Listrus  niveicanthus,  new  species. — Form  oblong-oval,  slightly  robust 
and  moderately  convex.  Color  black,  with  a  feeble  aeneous  tinge 
anteriorly;  mouth-parts  pale;  antennae  and  tarsi  rufous;  tibiae  toward 
apex  more  or  less  rufous.  First  antennal  joint  black  as  usual. 

Pubescence  moderately  short,  rather  fine,  recumbent,  somewhat  sparse 
and  plumbeo-cinereous  in  color ;  a  tuft  of  dense  snowy-white  hairs  at 
anterior  canthus  of  each  eye.  The  areas  of  dark  brown  hairs  are 
arranged  as  follows :  Basal  maculae  more  or  less  obsolete  or  small  when 
present,  the  humeral  frequently  absent  when  a  parascutellar  is  present  on 
each  elytron ;  post-basal  macula  not  large  but  variable  in  size,  tending  to 
extend  toward  the  humeral ;  submedian  very  narrow  and  zig-zag,  often 
interrupted  at  suture  and  middle  of  each  elytron ;  subapical  usually  inter- 
rupted forming  a  transverse  row  of  four  maculae,  sometimes  nearly  entire 
or  the  sutural  two  may  coalesce  to  form  a  small  rhomboidal  macula  on 
the  suture;  apical  maculae  obsolete  or  small  and  somewhat  transverse. 
Pronotal  central  figure  usually  dissolved  into  four  maculae,  two  anterior 
and  two  posterior,  which  with  those  from  the  broken  lateral  vittae,  make 
a  circle  of  six  small  maculae;  in  some  instances  the  constricted  figure 
appears.  The  pubescence  is  longer  on  the  under  surface  of  the  body, 
snowy-white  and  dense  on  the  prosternum  and  sternal  side-pieces,  duller 
in  color  on  the  abdomen  and  not  quite  as  dense. 

Head  somewhat  transverse,  distinctly  wider  than  the  pronotal  apex, 
eyes  moderately  large  and  prominent ;.  front  broadly  impressed,  a  small 
and  somewhat  glabrous  convexity  just  behind  the  base  of  the  epistoma ; 
surface  densely  indentato-punctate.  Antennae  moderate  in  length  and 
nearly  similar  in  the  sexes,  moderately  compressed;  second  joint  oval,  a 
little  longer  than  wide;  third  equal  in  length  to  the  second,  obconico- 
cylindrical  in  form,  a  little  less  than  twice  as  long  as  wide ;  fourth  and 
fifth  noticeably  compressed,  fourth  slightly  elongate  and  triangular, 
moderately  prominent  anteriorly,  apical  margin  oblique;  fifth  about  as 
wide  as  the  ninth,  distinctly  wider  than  the  joints  between  the  first  and 
ninth,  slightly  elongate,  about  a  half  longer  than  wide,  angulate  anteriorly, 
apical  margin  oblique;  sixth,  seventh  and  eighth  triangular,  subequal  in 
length,  just  a  little  longer  than  wide  and  slightly  angulate  anteriorly; 
ninth  and  tenth  triangular,  anterior  margins  moderately  arcuate,  subequal 
in  width  with  the  eleventh,  the  latter  less  than  a  half  longer  than  wide, 
oval,  slightly  narrower  at  apex. 


168      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Pronotum  transverse,  about  a  fourth  wider  than  long,  almost  evenly 
convex;  apex  broadly  arcuate;  sides  broadly  and  moderately  strongly 
arcuate,  widest  a  little  behind  the  middle,  arcuately  convergent  anteriorly, 
angles  broadly  rounded,  the  basal  sometimes  slightly  obtuse ;  base  broadly 
arcuate,  feebly  sinuate  laterally  within  the  angles  when  obtuse ;  disk  dull 
in  lustre  and  densely  indentato-punctate. 

Elytra  oblong,  about  twice  as  long  as  wide ;  apex  rather  parabolically 
rounded ;  not  deeply  punctate,  punctures  separated  by  a  distance  equal  to 
one  or  two  times  their  diameter,  surface  more  or  less  transversely  and 
feebly  rugose,  punctures  finer  toward  apex. 

Abdomen  finely  and  very  densely  granulato-punctate ;  pubescence 
almost  hiding  the  body  surface ;  fifth  ventral  segment  modified  on  the  disk 
in  the  male. 

Legs  moderate  in  length  and  stoutness ;  metafemora  rather  strongly 
arcuate  dorsally  in  basal  third,  rather  stout  and  subparallel,  i.  e.,  less 
narrowed  at  base. 

Male. — Slightly  narrower,  subparallel,  antennae  slightly  heavier; 
fifth  ventral  segment  distinctly  sinuate,  adjacent  surface  impressed  and 
asperate,  impressed  area  invested  with  black  hairs  which  are  not  noticeably 
bristling,  pubescence  unmodified  beyond  margin  of  the  impression,  some- 
times brownish,  especially  on  the  genital  segment,  tactile  hairs  rather  long 
and  somewhat  numerous. 

Female. — Slightly  broader,  a  little  more  ovate ;  antennae  a  little  more 
slender,  anterior  margin  rather  less  serrate.  Metafemora  rather  more 
parallel ;  fifth  ventral  broadly  rounded  at  apex. 

Measurements. — Length  (Types),  3.2  mm.;  width,  1.2-1.3  mm. 

Holotype,  male,  in  my  own  collection ;  allotype,  female,  in  that  of  the 
California  Academy  of  Sciences ;  paratypes  in  both  collections. 

Type  locality. — Bartlett  Springs,  Lake  County,  California.  Collected 
in  June  by  Dr.  A.  Fenyes. 

Distribution. — Besides  the  typical  phase  taken  at  Bartlett  Springs,  I 
have  eight  specimens  from  Pasadena,  in  Southern  California.  These  are 
smaller  with  the  apical  joints  of  the  antennae  somewhat  darker,  and  the 
central  pronotal  constricted  macula  is  represented  by  a  rounded  macula 
corresponding  to  the  basal  lobe  of  the  "hourglass"  spot.  In  the  males 
the  impressed  area  of  the  fifth  ventral  is  quite  glabrous  and  the 
pubescence  is  longer  at  the  angles.  In  some  Listri  there  is  an  apparent 
sixth  segment,  but  it  is  exceedingly  difficult  to  determine  the  true  structure 
in  dried  specimens,  especially  when  they  are  few  in  number.  Careful 
dissection  is  necessary. 

From  reading  Casey's  description  of  Listrus  tritus  Casey,  one  would 
refer  niveicanthus  to  that  species  as  regards  maculation.  In  tritus  the 


DESCRIPTION  OF  NEW  SPECIES  169 

fifth  ventral  segment  in  the  male  is  truncate  at  apex  and  unmodified  on 
the  disk;  in  Udells  Casey  it  is  likewise  truncate  and  unmodified,  the 
pubescence  being  of  the  usual  character.  Extricatus  Casey  is  recognized 
and  properly  placed  from  material  taken  at  San  Diego  and  Pasadena,  in 
this  species  the  antennae  are  stouter  and  incrassate,  and  blackish  in  color, 
except  the  second  joint  which  is  rufous  or  testaceous.  The  above  variant 
of  niveicanthiw  taken  at  Pasadena  may  be  known  as  tincticornis,  new  sub- 
species. 

Listrus  elegantulus,  new  species. — Form  oblong-oval  to  oblong-ovate, 
rather  strongly  convex.  Color  black,  a  feeble  purpureo-aeneous  lustre, 
especially  on  the  head  and  pronotum ;  mouth  pale,  second  joint  of  the 
antennae  rufo-testaceous,  succeeding  joints  more  or  less  rufo-piceous, 
apical  joints  usually  blackish ;  tibiae  dark  rufo-piceous,  tarsi  more  or  less 
rufous. 

Pubescence  moderately  long,  abundant  and  cinereous  in  color;  dark 
areas  of  dark  brownish  hairs  arranged  as  follows:  Elytral  maculation 
conspicuous,  basal  fascia  rather  broad,  usually  narrowly  interrupted  at 
middle  by  the  albescent  scutellum ;  post-basal  macula  small  at  middle  of 
each  elytron ;  median  fascia  quite  broad,  edges  irregular,  extending  some- 
what along  the  suture  anteriorly ;  subapical  fascia  narrower,  with  edges 
zig-zag,  widening  more  or  less  on  the  suture  to  form  a  subrhomboidal 
figure,  and  constricted  at  middle  of  each  elytron  almost  to  the  point  of 
division ;  apical  maculae  rather  large,  but  variable  as  to  size.  Pronotal 
central  macula  usually  distinctly  constricted  at  middle,  basal  lobe  rather 
larger  than  the  apical,  reentering  lines  not  evident;  lateral  vittae  more 
or  less  obsolete  or  divided  at  middle.  Central  area  of  the  head  with 
darkish  hairs. 

Head  somewhat  transverse  and  wider  than  the  pronotal  apex,  eyes 
large  and  prominent ;  front  more  or  less  broadly  impressed,  at  times 
apparently  bi-impressed,  a  feeble  median  convexity  behind  the  epistomal 
base ;  surface  densely  indentato-punctate ;  sometimes  the  intervals  are 
smooth  on  the  vertex  and  adjacent  front  where  the  punctures  are  rather 
coarse,  but  anteriorly  between  the  antennae  they  are  more  densely  placed. 
Antennae  rather  long  and  slender,  feebly  compressed  toward  base,  apical 
joints  feebly  incrassate;  second  joint  longer  than  wide  and  rather 
obconical ;  third  rather  slender  and  cylindrical,  about  as  long  as  the  fourth 
or  second ;  fourth  more  evidently  compressed,  obconico-subtriangular, 
slightly  prominent  anteriorly  toward  apex;  fifth  slightly  elongate,  sub- 
triangular,  apical  margin  rather  oblique,  anterior  border  angulate,  dis- 
tinctly wider  than  the  fourth  or  third,  about  equal  in  width  to  the  eighth ; 
sixth  and  seventh  just  a  little  narrower,  less  noticeably  compressed,  sub- 
triangular  and  very  slightly  longer  than  wide ;  eighth  triangular  in  outline 


170      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

and  not  noticeably  compressed ;  ninth  distinctly  wider  than  long  and  with 
the  tenth  and  eleventh  just  a  little  wider  than  the  fifth  ;  tenth  a  little  longer 
than  the  ninth,  subtriangular ;  eleventh  not  more  than  twice  as  long  as 
wide,  nearly  an  elongate  oval. 

Pronotum  about  a  fourth  wider  than  long,  evenly  convex,  more 
declivous  antero-laterally ;  apex  arcuato-truncate  in  circular  arc;  sides 
broadly  arcuate,  less  so  anteriorly  and  moderately  convergent  to  apex, 
serrules  short  and  more  or  less  acute,  fimbriae  rather  short  and  not  closely 
placed;  angles  broadly  rounded,  the  anterior  less  so;  base  broadly  and 
rather  evenly  arcuate,  scarcely  sinuate  laterally  within  the  angles;  disk 
in  the  central  area  rather  sparsely  punctured,  punctures  less  than  coarse, 
not  very  sharply  defined  although  deep,  intervals  rather  irregular  in  width 
and  from  two  to  four  times  wider  than  the  punctures,  not  noticeably 
indented,  lateral  thirds  distinctly  and  moderately  coarsely  and  densely 
indentato-punctate. 

Elytra  somewhat  oblong-oval,  sides  moderately  arcuate  although  sub- 
parallel,  about  twice  as  long  as  wide;  discal  punctures  not  coarse,  sepa- 
rated by  a  distance  equal  to  one  or  three  times  their  diameter,  finer  toward 
apex  and  more  widely  spaced,  surface  feebly  transversely  rugose. 

Abdomen  finely  and  rather  densely  punctate;  fifth  ventral  segment 
modified  on  the  disk  in  the  male. 

Legs  moderate  in  length  and  stoutness ;  tarsi  of  medium  length. 

Male. — Oblong-oval,  subparallel,  somewhat  robust  as  compared  to 
the  male  of  bifasciatus.  Antennae  just  a  little  stouter  than  in  the  female. 
Fifth  ventral  segment  distinctly  emarginate,  emargination  arcuate  and 
moderately  deep,  lateral  angles  prominent,  impressed  surface  of  the  seg- 
ment subasperate,-  pubescence  longer  and  blackish  to  brownish.  Meta- 
femora  moderately  stout  and  subfusiform. 

Female. — Oblong-ovate,  rather  more  convex  posteriorly,  sides  a  little 
more  arcuate ;  pronotal  sides  more  strongly  arcuate  posteriorly.  Antennae 
slightly  shorter  and  a  little  more  slender;  fifth  ventral  rather  broadly 
rounded  at  apex ;  abdominal  apex  with  blackish  pubescence. 

Measurements. — Length  (Types),  3-3.3  mm.;  width,  l-1.2mm. 

Holotype,  male,  allotype,  female,  in  my  own  collection ;  paratypes  in 
that  of  the  California  Academy  of  Sciences  and  that  of  Mr.  F.  W.  Nunen- 
macher. 

Type  locality. — Willow  Creek,  Humboldt  County,  California.  Col- 
lected by  Mr.  Nunenmacher  on  May  20,  1911,  and  myself  on  June  13,  1916. 

Distribution. — California  (Humboldt  County;  Siskiyou  County, 
June  2,  1911 ;  Mokelumne  Hill,  Calaveras  County,  April). 

This  elegant  species  in  its  oblong-ovate  or  oblong-oval  form,  rather 
wide  elytral  fasciae,  quite  slender  antennae  which  are  very  nearly  similar 


DESCRIPTION  OF  NEW   SPECIES  171 

in  the  sexes,  and  modified  fifth  ventral  in  the  male  and  cinereous 
pubescence,  gives  a  facies  notably  different  from  the  other  species 
described  in  the  present  paper  and  also  those  previously  described  by 
Col.  Casey. 

Elegantulus  is  apparently  related  to  interruptus  Lee.  According  to 
Casey,  in  interruptus  the  dark  pubescence  of  the  elytra  is  arranged  in  spots, 
and  he  distinctly  states  that  the  interstitial  spaces  of  the  head  and 
pronotum  are  polished  and  without  reticulation.  In  interruptus  the  fifth 
ventral  segment  in  the  male  is  strongly  sinuato-truncate  at  apex,  flattened 
on  the  disk  and  clothed  with  long  erect  and  bristling  hairs,  which  become 
black  in  apical  half  of  the  segment.  In  elegantulus  the  hairs  are  not  very 
bristly,  although  rather  long  and  brownish  to  black  in  color ;  the  apex  of 
the  fifth  ventral  in  the  male  is  distinctly  emarginate,  with  the  lateral  angles 
quite  prominent. 

Interruptus  is  distributed  from  Nebraska  to  California,  where  it  does 
not  descend  the  western  slope  of  the  Sierras  in  California,  according  to 
Col.  Casey. 

Listrus  fulvipilosus,  new  species. — Form  subparallel  to  elongate-oval 
and  convex.  Color  black,  with  a  feeble  bluish  metallic  lustre  and  rather 
shining ;  antennae  more  or  less  rufo-piceous ;  tibiae  dark  piceous,  tarsi 
rufous  to  rufo-piceous. 

Pubescence  rather  short,  sparse  and  somewhat  coarse,  for  the  greater 
part  fulvous  in  color,  varying  in  some  specimens  to  plumbeo-cinereous. 
The  darker  hairs  are  arranged  in  an  obscure  elytral  pattern  as  follows : 
A  small  or  moderate  humeral  and  a  parascutellar  macula  on  each  elytron ; 
a  post-basal  at  middle  of  each ;  a  narrow  zig-zag  fascia  at  middle,  which 
may  be  dissolved  into  a  transverse  row  of  narrow  irregular  maculae ; 
subapical  transverse  row  of  more  or  less  distinct  maculae,  and  one  on  each 
elytron  at  apex.  In  some  specimens  the  markings  are  quite  distinct  when 
viewed  longitudinally  from  behind.  Pronotal  central  figure  indistinct,  or 
evidently  constricted  at  middle  with  the  lobes  divided  longitudinally  by 
a  median  line  of  pale  hairs ;  lateral  vittae  more  or  less  dissolved  into  two 
maculae.  Elytral  pattern  similar  to  that  observed  in  niveicanthus  and 
its  subspecies  tincticornis.  Hairs  of  the  under  surface  of  the  body  rather 
long. 

Head  about  as  wide  as  long,  quite  strongly  and  broadly  bi-impressed 
between  the  eyes  and  antennae,  impressions  separated  by  a  feeble  median 
convexity  that  is  more  or  less  subglabrous  toward  the  epistomal  base,  the 
latter  more  or  less  transversely  rugulose ;  surface  rather  coarsely  and 
densely  indentato-punctate,  slightly  strigose  against  the  eyes.  Antennae 
rather  slender  and  moderately  dissimilar  in  the  sexes,  sometimes  pale  in 
color  and  apparently  extending  to  the  pronotal  base. 


172      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Pronotum  transverse,  about  a  fourth  wider  than  long,  widest  at  basal 
third ;  sides  broadly  arcuate  in  basal  half  and  less  so  anteriorly,  con- 
verging moderately  to  apex,  serrules  small  and  subacute,  fimbriae 
moderate  in  length ;  apex  broadly  arcuato-truncate,  angles  obtusely  and 
rather  broadly  rounded;  base  broadly  arcuate  in  middle  six-eighths, 
sinuate  within  the  angles,  the  latter  subdentate  and  obtuse;  disk  moder- 
ately arcuate  from  side  to  side,  more  strongly  so  anteriorly,  more  or  less 
coarsely  indentato-punctate,  densely  so  laterally,  less  so  in  the  central  area 
where  the  indentations  are  rather  shallow. 

Elytra  slightly  more  than  twice  as  long  as  wide,  at  base  about  equal  to 
the  width  of  the  pronotum ;  rather  coarsely  punctate,  punctures  separated 
by  a  distance  equal  to  one-half  to  three  times  their  diameter,  surface  finely 
and  rather  obsoletely  rugulose;  punctures  much  finer  toward  apex,  the 
latter  subserrulate  on  the  edge,  sutural  angles  obtusely  but  not  strongly 
rounded. 

Abdomen  finely,  rather  densely  punctate;  fifth  ventral  segment 
different  in  the  sexes,  being  modified  on  the  disk  in  the  male. 

Legs  moderate  in  length  and  stoutness ;  metafemora  moderate  in 
stoutness  with  their  dorsal  outline  distinctly  arcuate. 

Male. — Parallel  and  slightly  oblong.  Antennae  with  the  second  joint 
oval,  slightly  narrowed  at  the  base,  about  a  third  longer  than  wide,  quite 
equal  in  length  to  the  third ;  the  latter  cylindrical,  twice  as  long  as  wide, 
comparatively  narrow ;  fourth  obconical,  feebly  prominent  anteriorly, 
about  a  half  wider  than  long,  subequal  in  length  to  the  third ;  fifth  rather 
elongate,  subtriangular,  about  a  third  longer  than  the  width  of  the  apex, 
apical  margin  transverse,  as  wide  as  the  seventh,  distinctly  wider  than  the 
sixth,  fourth,  third  or  second,  moderately  prominent  anteriorly ;  sixth  and 
eighth  triangular,  about  as  long  as  wide  and  subequal ;  seventh  almost 
triangular,  longer  than  wide,  sides  slightly  arcuate ;  ninth  wider,  tri- 
angular, about  as  long  as  wide ;  tenth  subtriangular,  less  narrowed  toward 
base,  sides  noticeably  arcuate,  very  slightly  longer  than  wide ;  eleventh 
elongate  oval,  feebly  narrowed  apically,  almost  twice  as  long  as  wide. 
Fifth  ventral  segment  modified ;  broadly  and  rather  deeply  emarginate, 
emargination  rounded,  adjacent  surface  of  the  segment  rather  glabrous 
and  shining,  the  impressed  area  semilunar  in  outline,  surface  beyond  sub- 
asperate,  set  with  rather  long  but  not  dense  blackish  hairs  which  extend 
somewhat  on  to  the  apex  of  the  fourth  segment ;  hairs  rather  long  at  the 
angles  of  the  emargination,  the  latter  rather  obtuse.  Modified  area 
blackish. 

Female. — Subparallel,  sides  of  the  body  slightly  arcuate.  Antennae 
more  slender,  joints  proportioned  as  in  the  male;  the  fourth  rather  sub- 
triangular;  fifth  slightly  narrower,  less  prominent  anteriorly,  apical 


DESCRIPTION  OF  NEW   SPECIES  173 

margin  rather  oblique,  obtusely  angulate  anteriorly ;  tenth  noticeably  less 
stout,  about  as  long  as  wide,  quite  triangular ;  eleventh  obovate,  shorter, 
about  a  half  longer  than  wide.  Fifth  ventral  broadly  and  evenly  arcuate 
at  apex,  somewhat  broadly  flattened,  margin  slightly  deflexed. 

Measurements. — Length  (Types),  3.0-3.5  mm.;  width,  1.1-1.3  mm. 

Holotype,  male,  and  allotype,  female,  in  my  own  collection  ;  paratypes 
in  that  of  Mr.  F.  W.  Nunenmacher,  who  collected  the  material  on  June  4, 
1913. 

Type  locality. — Lassen  County,  California. 

Twenty  specimens  studied.  Fulvipilosus  resembles  amplicollis  Casey, 
from  which  it  differs  in  the  densely  punctured  pronotum,  male  sexual 
characters  and  more  or  less  fulvous  pubescence. 

Listrus  cervicalis,  new  species. — Form  oblong-oval  and  moderately 
convex.  Color  black,  with  a  slight  aeneous  lustre,  chiefly  on  the  head  and 
pronotum  ;  antennae  more  or  less  piceous,  second  joint  rufo-piceous ;  tibiae 
and  tarsi  piceous. 

Pubescence  moderate  in  length,  abundant  and  plumbeo-cinereous  in 
color,  longer  on  the  under  surface  of  the  body,  cinereous  on  the  sterna 
and  sternal  side-pieces,  slightly  darker  on  the  abdomen.  Brownish  black 
hairs  of  the  elytra  are  arranged  in  rather  large  maculae  as  follows:  A 
narrow  and  elongate  humeral  and  a  rounded  parascutellar  macula ;  post- 
basal  macula  at  middle  of  each  with  a  tendency  to  diffuse  toward  the 
humeral ;  a  submedian  rather  wide  and  more  or  less  zig-zag  fascia  which 
may  or  may  not  be  interrupted  at  the  suture  or  at  middle  of  each  elytron ; 
subapical  transverse  row  of  four  rather  large  maculae ;  apical  maculae  of 
good  size.  There  is  a  tendency  for  the  maculae  to  coalesce  transversely  to 
form  five  transverse  dark  fasciae.  Pronotal  central  figure  constricted  and 
more  or  less  divided  by  a  narrow  median  line  of  pale  hairs  into  four 
maculae,  lateral  vittae  broken  into  two  distinct  maculae;  hairs  of  the 
central  area  of  the  frons  more  or  less  dark.  Hairs  of  the  pronotum  are 
rather  stiff,  closely  recumbent  and  somewhat  abundant. 

Head  rather  large,  about  as  long  as  wide,  eyes  moderately  prominent 
and  evenly  convex.  Front  broadly  impressed,  a  slight  median  convexity 
near  the  epistomal  base,  punctures  moderate  in  size  and  rather  sparse, 
denser  at  the  periphery,  intervals  more  or  less  smooth  and  more  or  less 
feebly  punctulate  or  subindentate ;  muzzle  short  and  rather  broad,  margin 
evenly  arcuate  from  side  to  side.  Antennae  quite  long,  somewhat  slender, 
and  only  slightly  compressed,  feebly  incrassate  and  similar  in  the  sexes ; 
second  joint  oval  narrowing  somewhat  toward  base ;  third  cylindro- 
obconical  and  as  long  as  the  fourth,  the  latter  wider  and  triangulo- 
obconical ;  fifth  wider,  about  as  wide  as  the  ninth,  triangular  and  just  a 
little  longer  than  wide,  quite  prominent  anteriorly ;  sixth  to  the  eighth 


174      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

subequal  in  length  and  size,  subtriangular,  almost  circular  in  transverse 
section ;  ninth  and  tenth  stouter,  a  little  longer,  subtriangular  and  circular 
in  section ;  eleventh  obovate,  rather  stout,  about  a  half  longer  than  wide. 

Pronotum  about  a  fourth  wider  than  long,  slightly  transversely 
oblong;  apex  wide,  moderately  broadly  arcuate  in  circular  arc;  apical 
angles  rather  obtusely  rounded;  sides  quite  broadly  arcuate,  converging 
very  moderately  anteriorly,  more  or  less  feebly  sinuate  before  the  basal 
angles  (see  sexes),  serrules  moderate  in  size  and  rather  irregular, 
fimbriae  moderate  in  length ;  base  broadly  and  rather  strongly  arcuate, 
feebly  sinuate  laterally,  angles  apparently  rounded  although  subobtuse 
from  a  rather  broad  and  reflexed  denticle ;  disk  quite  strongly  convex, 
rather  strongly  declivous  antero-laterally,  punctures  rather  large,  some- 
what widely  spaced  in  the  central  area,  intervals  flat,  more  or  less 
indentated  toward  base,  densely  indentato-reticulato-punctate  laterally. 

Elytra  scarcely  twice  as  long  as  wide,  oblong,  apex  rather  broadly 
and  evenly  rounded ;  sides  parallel  and  slightly  arcuate ;  punctures  rather 
coarse,  separated  by  a  distance  equal  to  two  or  three  times  their  diameter ; 
surface  smooth,  punctures  finer  toward  apex. 

Abdomen  finely  and  more  or  less  densely  ruguloso-punctulate ;  fifth 
ventral  segment  modified  on  the  disk  in  the  male. 

Legs  moderately  stout  and  a  little  longer  than  usual. 

Male. — Slightly  narrower  and  a  little  more  parallel.  Pronotal  sides 
rather  less  arcuate,  as  a  rule  slightly  sinuate  before  the  basal  angles, 
serrules  a  little  coarser,  basal  angles  more  obtuse.  Fifth  ventral  broadly 
and  somewhat  deeply  sinuate,  lateral  angles  rather  prominent  and  narrowly 
rounded,  adjacent  surface  distinctly  impressed  and  more  or  less  glabrous, 
margin  of  impression  asperate  and  set  with  rather  stout  black  hairs  which 
may  extend  forward  on  to  the  fourth  segment  for  a  short  distance ;  termi- 
nal segment  of  the  abdomen  set  with  shorter  black  hairs. 

Female. — A  little  broader  and  more  arcuate  at  the  sides.  Pronotal 
sides,  as  a  rule  more  arcuate,  scarcely  sinuate  before  the  basal  angles,  the 
latter  more  broadly  rounded.  Fifth  ventral  not  very  broad  at  apex,  sides 
converging  more  strongly,  tip  not  very  strongly  rounded,  margin  narrowly 
deflexed  and  asperate,  surface  of  the  segment  feebly  impressed. 

Measurements. — Length  (Types),  3.3-3.5  mm.;  width,  1.2-1.3  mm. 

Holotype,  male,  and  allotype,  female,  both  in  the  collection  of  the 
California  Academy  of  Sciences  ;  paratypes  in  the  Academy's  and  my  own 
collection. 

Type  locality. — Huntington  Lake,  Fresno  County,  California,  at  an 
elevation  of  7000  feet.  Beaten  from  fir  trees.  A  large  series  were 
secured  by  Mr.  E.  P.  Van  Duzee  and  myself  during  July,  1919. 


SYNOPSIS  OF  SPECIES  OF  LISTRUS  175 

Ceruicalis  at  first  sight  resembles  montanus  Casey ;  in  the  former  the 
maculation  is  more  distinct,  pronotum  broader  and  more  convex  and  the 
fifth  ventral  segment  is  modified  in  the  male,  besides  there  is  a  tendency 
for  the  elytral  maculae  to  unite  to  form  five  transverse  dark  fasciae,  which 
are  frequently  in  evidence  and  unique. 

SYNOPSIS  OF  THE  SPECIES  OF  LISTRUS. 

The  species  of  Listrus  may  be  divided  into  two  Sections  as  follows : 
Fifth    ventral    abdominal    segment    in    the    male,    truncate    or    sinuato- 

truncate   at   apex Section    I 

Fifth  ventral  more  deeply  sinuate  and  modified  on  the  disk  in  the  male Section  II 

The  species  can  be  arranged  in  the  following  Groups : 

SECTION  I. 

Elytral  apices  different  in  the  sexes Definitus  Group 

Elytral  apices  similar  in  the  sexes 1 

1.  Eyes  very  prominent  and  more  strongly  convex  in  anterior  two-thirds; 

head  broad,  male Cephalicus  Group 

— .     Eyes  moderately  prominent  and  evenly  convex 2 

2.  Elytral  pubescence  unicolorous Senilis  Group 

— .     Elytra  ornate  with  a  pattern  of  blackish  or  brownish-black  hairs 3 

3.  Elytral  pubescence  very  short,  pattern  very  obscure  from  sparseness  of 

pale  hairs  Punctatus  Group 

— .    Elytral  pubescence  longer  and  more  abundant,  pattern  distinct 4 

4.  Elytra  with  three  rather  wide  transverse  black  fasciae,  the  basal  of  which 

may  be  more  or  less  interrupted Annulatus  Group 

— .     Elytral  fasciae  more  or  less  modified  or  broken  up 5 

5.  Species   with  legs   more  or   less   piceous  or  rufo-piceous,   femora  never 

pale  DifHcilis  Group 

— .  Species  usually  scarcely  over  2.5  mm.  in  length,  except  luteipes  (3  mm.)  ; 
legs  entirely  pale;  basal  and  post-basal  fasciae  broken  up  into  variable 
small  maculae  Luteipes  Group 

SECTION  II. 

Metatrochanters  triangular Trochantericus  Group 

Metatrochanters  of  the  usual  form,  suboval 1 

1.  Pubescence  short  and  sparse,  elytral  pattern  obscure  but  discernible 

Liebecki  Group 

— .     Pubescence  longer,  elytral  pattern  distinct 2 

2.  Elytral  maculation  tending  to  form  fasciae  and  nearly  constant  in  charac- 

ter   --  Interruptus  Group 

— .     Elytral  maculation  consisting  of  isolated  maculae  tending  to  obsolescence 

Salicis   Group 

DEFINITUS  GROUP. 

Elytra  with  a  wide  subapical  black  fascia;  apices  acuminately  produced  in  the 
female  and  rounded  as  usual  in  the  male;  pronotum  longer  than  wide; 
legs  entirely  pale 1.  definitus  Fall 


176      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 
CEPHALICUS   GROUP. 

Head  broad,  eyes  very  prominent  and  strongly  convex  anteriorly,  forming  a 
straight  and  posteriorly  convergent  line  with  the  temporal  region ;  elytral 
maculation  obscure ;  legs  and  antennae  stout 2.  cephalicus,  n.  sp. 

SENILIS  GROUP. 

Body  densely  and  uniformly  clothed  with  coarse  and  dense  cinereous  hairs 

- 3.     senilis  Lee. 

Body  more  sparsely  clothed  with  finer  and  less  cinereous  pubescence 1 

1.  Tenth  antennal  joint  almost  as  long  as  wide;  male  elongate  and  parallel; 

antennae  distinctly  longer  than  the  pronotum 4.     uniformis  Casey 

— .    Tenth  antennal  joint  transverse 2 

2.  Fifth   ventral  abdominal  segment  truncate   at  apex,  male;   pronotal  disk 

broadly,  evenly  convex 5.    clavicornis  Casey 

— .    Fifth    ventral    longer,    arcuato-truncate    at    apex,    male;    pronotal    disk 

arcuately  declivous  antero-laterally 6.    coloradensis,  n.  sp. 

PUNCTATUS   GROUP. 

Basal  angles  of  the  pronotum  distinctly  obtuse  and  angulate;  sixth  and  eighth 

antennal  joints  distinctly  smaller  than  the  contiguous  joints 

7.  obscurellus  Lee. 

Basal  angles  of  pronotum  broadly  rounded  and  quite  continuously  so  with  the 

sides  and  base 1 

1.  Smaller  species  and  not  elongate;  antennae  subclavate,  joints  four  to  eight 

scarcely  longer  than  wide;  frons  and  central  area  of  pronotum  sparsely 

and  distinctly  punctate;  fimbriae  very  short 8.  punctatus  Motsch. 

— .  Larger  species;  antennae  not  clavate,  long  and  slender,  joints  distinctly 

elongate ;    pronotum    more    transverse ;    fimbriae    moderate    in    length ; 

pubescence  longer;  elytral  pattern  discernible 9.    dilutus,  n.  sp. 

ANNULATUS   GROUP. 

Elytra  with  three  transverse  fasciae,  the  median  widest,  tending  to  diffuse  along 

the  suture ;  head  and  pronotum  noticeably  narrower  than  the  elytral  base ; 

central  area  of  pronotum  very  sparsely  punctate 10.  martini,  n.  sp. 

Elytra  with  three  almost  even  transverse  dark  fasciae ;  surface  somewhat  dull ; 

fourth  and  fifth  antennal  joints  triangular 1 

1.  Legs  piceous  or  black;  form  stouter  and  oblong  (male),  or  oblong-ovate 

(female)  11.  annulatus  Casey 

— .  Legs  rufous;  form  feebly  elongate,  oblong-oval 12.  rubripes  Casey 

DIFFICILIS  GROUP. 

Pronotum  strongly  and  closely  indentato-punctate  in  the  central  area 1 

Pronotum  not  closely  punctate  in  the  central  area  but  with  the  intervals  flat 

and  more  or  less  smooth 7 

1.  Third  antennal  joint  in  the  male  with  a  moderately  stout  chitinous  seta 

near  the  antero-apical  angle ;  male  parallel  and  elongate,  antennae  heavy ; 

female  elongate  ovate,  antennae  moderately  slender 13.     maculosus  Casey 

— .     Third  antennal  joint  in  the  male  without  a  stouter  seta 2 

2.  Third  antennal  joint  elongately  triangular;  elytral  maculae  more  or  less 

rounded  14.    montanus  Casey 

— .    Third  antennal  joint  more  or  less  cylindrical 3 


SYNOPSIS  OF  SPECIES  OF  LISTRUS  177 

3.  Dark  elytral   maculae   small   and   few   in   number,   pubescence   otherwise 

cinereous  15.    confusus  Casey 

— .     Dark  elytral  maculae  well  developed 4 

4.  Tibiae  and  tarsi   clear    rufous ;   lustre  more   or  less   cupreous ;    antennae 

similar   in   the    sexes,    fifth   joint   scarcely   wider    than   the   contiguous 

Joints 16.    extricatus  Casey 

— .     Tibiae  and  tarsi  more  obscure,  black  to  rufo-piceous  in  color 5 

5.  Elytra  with  submedian  and  subapical  fasciae 17.     difficilis  Lee. 

— .     Elytra  without  two  distinct  fasciae 6 

6.  Elytral  subapical  fascia  alone  distinct ;  submedian  resolved  into  two  strongly 

V-shaped  maculae ;  pubescence  dense  and  very  coarse 18.    fidelis  Casey 

— .     Elytral  dark  areas  in  large  maculae;  pubescence  rather  short,  sparse  and 

readily  removable 19.    subaeneus  Casey 

7.  Tarsi  and  distal  part  of  tibiae  rufous 8 

— .    Tarsi  and  tibiae  more  or  less  piceous 9 

8.  Submedian  elytral  fascia  zig-zag,  narrow,  V-shaped  on  each  elytron;  sub- 

apical    forming   a    moderate    rhomboidal   macula   on    the   suture,    apical 

maculae  rather  large ;  tarsi  and  tibiae  rufous _ 20.    vestitus,  n.  sp. 

— .     Submedian  and  subapical  fasciae  transverse,  the  latter  rather  broad;  tarsi 

and  distal  half  of  tibiae  rufous  or  paler 21.    incertus  Casey 

9.  Size  larger.     Pubescence  noticeably  long;  elytral  maculation  complex,  sub- 

apical  fasciae  broken  into  four  rather  large  maculae,  the  two  median  of 

which  may  form  a  subcordate  macula  on  the  suture 10 

— .     Size  smaller;  elytral  pattern  simpler 12 

10.  Form  different  in  the  sexes;  male  rather  narrow,  parallel  and  elongate; 

female  oblong-ovate ;  second  antennal  joint  a  half  longer  than  wide,  third 
obconic,  fourth  distinctly  triangular;  pubescnce  quite  abundant  on  ab- 
dominal apex  22.  bifasciatus,  n.  sp. 

— .  Form  somewhat  similar  in  the  sexes ;  male  oblong-oval ;  female  oblong- 
ovate;  antennae  similar  in  the  sexes;  third  and  fourth  antennal  joints 
elongate  and  subequal  in  length 11 

11.  Fifth  antennal  joint  distinctly  longer  than  wide  at  apex.     Surface  with  a 

purplish   metallic   reflection;   fifth  ventral  segment  broadly  truncate   at 

apex;  pale  elytral  fasciae  narrow  and  distinctly  zig-zag 23.    pardalis  Casey 

— .  Fifth  antennal  joint  about  as  long  as  wide  at  apex.  Surface  with  a  bluish- 
metallic  reflection;  fifth  ventral  broadly  and  feebly  sinuato-truncate ;  pale 

elytral  areas  broader  about  the  more  or  less  round  maculae 

24.     ntotschulskii  Lee. 

12.  Elytra  distinctly  fasciate  with  dark  hairs 13 

— .     Elytra    not    fasciate,    but    with    discrete    maculae    that   are   more   or   less 

atrophic  16 

13.  Upper  surface  of  body  shining  and  polished;  surface  with  a  greenish-  or 

bluish-metallic    lustre    14 

— .     Upper  surface  more  or  less  dull  and  with  a  cupreous  or  violaceous  lustre....     15 

14.  Elytra  with  three  distinct  and  rather  broad  transverse  fasciae,  the  basal 

narrowly  interrupted  at  scutellum,  post-basal  maculae  obsolete ;  pronotum 

quite  strongly  convex  25.    simplex,  n.  sp. 

— .  Elytra  without  three  fasciae;  pubescence  short,  sparse,  maculation  rather 
obscure ;  punctures  at  middle  of  pronotum  sparse,  intervals  more  or  less 
punctulato-rugulose  _ 26.  interstitialis  Casey 


178      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

15.  Surface  lustre  above  cupreous  throughout ;  pubescence  rather  short,  macu- 

lation  less  distinct;  antennae  rather   slender  basally,  last  three  joints 

stouter  and  feebly  incrassate 27.    variegatus  Casey 

— .  Surface  lustre  cupreous  on  head  and  pronotum,  violaceous  or  darker  on 
elytra;  pubescence  longer,  maculation  distinct,  antennae  somewhat 
stouter  28.  incestus,  n.  sp. 

16.  Surface  lustre  slightly  brassy ;  pubescence  rather  long,  coarse  and  cinereous, 

except  for  a  few  dark  maculae  behind  middle  of  the  elytra,  of  which 
four  are  arranged  in  a  subapical  transverse  row 29.     tritus  Casey 

LUTEIPES   GROUP. 

Antennal  joints  (male)  from  the  third  to  the  ninth  distinctly  elongate ;  pub- 
escence rather  short  and  cinereous;  median  and  subapical  fasciae  well 
denned  and  moderately  narrow ;  legs  entirely  pale...v 30.  giffardi,  n.  sp. 

Antennal  joints  not  elongate;  elytral  pubescence  unevenly  distributed  toward 

base 1 

1.  Head  and  thorax  noticeably  small,  distinctly  narrower  than  the  elytral  base ; 

elytral  dark  fasciae  more  or  less  diffused;  legs  pale 31.    parvicollis,  n.  sp. 

— .     Head  larger,  pronotum  about  as  wide  as  the  elytral  base 2 

2.  Pronotum  longer,  central  area  rather  sparsely  and  distinctly  punctate,  in- 

tervals more  or  less  smooth;  fourth  antennal  joint  obconico-triangular ; 

legs  pale 32.    luteipes  Lee. 

— .    Pronotum  shorter,  transverse,  densely  indentato-punctate 3 

3.  Albo-cinereous    hairs    of    the    elytra    elongate-lanceolate    and    somewhat 

squamiform,  very  dense  in  a  narrow  transverse  fascia  at  apical  third; 

fourth  antennal  joint  triangular;  legs  bright  rufous 33.    famelicus  Casey 

— .     Cinereous  hairs  of  the  elytra  slender  as  usual 4 

4.  Legs  piceous ;  elytral  pubescence  unevenly  distributed  anterior  to  a  moder- 

ately wide  subapical  dark  fascia;  fourth  antennal  joint  obconical 

34.     ornatulus  Casey 

— .    Legs  rufous  ,. _ 5 

5.  Smaller  species  (1.9  mm.);  fourth  antennal  joint  feebly  triangular;  basal 

joint  pale;  elytra  with  a  dark  wide  median  and  a  narrower  subapical 
fascia,  and  two  rounded  maculae  on  each  toward  base....3S.  concurrens  Casey 
— .  Larger  species  (1.9-2.3  mm.)  ;  fourth  antennal  joint  triangular  (male),  or 
obconical  (female)  ;  elytra  with  an  even  transverse  subapical  fascia,  apex 
with  a  few  scattered  dark  hairs,  and  feeble  trace  of  small  brownish 
maculae  toward  base _ 36.  balteellus  Casey 

TROCHANTERICUS   GROUP. 

Pubescence  coarse,  cinereous,  rather  dense;  form  feebly  elongate;  median 
elytral  fasciae  narrow,  zig-zag  or  broken  up ;  pronotal  central  area  closely 
indentato-punctate.  Antennae  moderate  in  length,  slightly  compressed; 
third  joint  cylindrical;  fourth  obconico-triangular;  sixth,  seventh  and 
eighth  subequal  in  size  and  length;  fifth  serrate  as  usual  in  the 
male  ^ 37.  tro chant ericus,  n.  sp. 

LIEBECKI   GROUP. 

Antennae  long  and  rather  stout;  joints  three,  four  and  five  elongate;  fourth 
quite  triangular;  third  rather  slender,  slightly  more  than  twice  as  long 
as  wide  (male) 38.  liebecki,  n.  sp. 


SYNOPSIS  OF  SPECIES  OF  LISTRUS  179 

Antennae  notably  stout;  joints  sixth  to  the  ninth  subequal  in  size  and  form; 
fourth  distinctly  triangular  and  scarcely  longer  than  wide;  third  rather 
short,  not  twice  as  long  as  wide  (male).  More  slender  and  subclavate 
in  the  female 39.  olympianus,  n.  sp. 

SALICIS  GROUP. 

Central  area  of  the  pronotum  not  indentato-punctate,  intervals  flat;  smaller 
species,  lustre  aeneous ;  antennae  and  legs  clear  rufous ;  antennal  joints 
six  to  nine  not  elongate 40.  occidens,  n.  sp. 

Central  area  of  the  pronotum  densely  indentato-punctate 1 

1.  Antennae  moderately  stout  and  blackish  in  color;  lustre  dark,  maculation 

well  developed;  fourth  antennal  joint  triangular;  fifth  joint  serrate  and 
about  as  wide  as  the  ninth  (male)  ;  pubescence  of  the  fifth  ventral  seg- 
ment scarcely  modified  in  the  male;  antennae  moderately  dissimilar  in 

the  sexes 41.    angulatus,  n.  sp. 

— .    Antennae  rufous  or  ruf o-piceous  in  color,  lustre  more  or  less  aeneous ; 

tibiae  and  tarsi  more  or  less  rufo-piceous 2 

2.  Fourth  antennal  joint  obconico-triangular ;  fifth  joint  scarcely  wider  than 

the  sixth;  joints  three  to  nine  moderately  elongate  in  the  male;  antennae 

moderately  dissimilar  in  the  sexes „ 42.    salicis,  n.  sp. 

— .  Fourth  antennal  joint  not  strongly  triangular;  joints  three,  four  and  five 
elongate,  six,  seven  and  eight  less  so;  fifth  not  strongly  serrate;  antennae 

more  slender  in  the  female 43.    niveicanthus,  n.  sp. 

Form  smaller,  apical  joints  of  antennae  darker iincticornis  n.  subsp. 

INTERRUPTUS   GROUP. 

Interstitial  spaces  of  the  central  area  of  the  pronotal  disk,  flat  and  smooth ; 

spaces  variable  in  width 1 

Interstitial    spaces    more    or    less    indentate;    surface    more    or    less    densely 

indentato-punctate 5 

1.  Pubescence  short,  very  sparse;  maculation  scarcely  discernible;  antennae 

slender,  more  or  less  rufo-piceous,  fifth  joint  scarcely  wider  than  the 

following  two  or  three  joints 44.    amplicollis  Casey 

— .     Pubescence  longer  and  denser;  maculation  distinct „.      2 

2.  Lustre  more  or  less  aeneous;  pubescence  moderately  long;  antennae  rufo- 

piceous  3 

— .     Lustre  dark,  bluish  or  greenish  metallic;  pubescence  shorter 4 

3.  Maculation  consisting  of  rounded  maculae  on  the  elytra;  fifth  ventral  of 

the  male  broadly  and  strongly  sinuato-truncate  at  apex,  flattened  on  the 
disk  and  clothed  with  long  erect  and  bristling  hairs  which  become  black 

in  apical  half 45.    interruptus  Lee. 

— .  Maculation  consisting  of  three  moderately  wide  fasciae,  median  and  sub- 
apical  with  irregular  edges;  fifth  ventral  clothed  with  brownish  gray 
hairs  on  the  flattened  area < 46.  elegantulus,  n.  sp. 

4.  Lustre  with  a  dark  steel-blue  metallic  reflection;  legs  and  antennae  black 

throughout ;  pubescence  very  short,  decumbent  and  coarse,  a  median  and 
subapical  fascia  with  basal,  post-basal  and  apical  maculae,  or  with  fasciae 

broken  into  maculae., 47.     rotundicollis  Casey 

— .  Lustre  with  a  virido-aeneous-  reflection;  legs  and  antennae  more  or  less 
picescent,  second  joint  paler;  pubescence  rather  short,  dark  areas  large, 
two  wide  and  straight  transverse  fasciae  on  the  elytra  behind  the 


180      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRION1DAE 

middle  48.    canescens  Mann. 

5.  Antennae  rufo-testaceous,  tibiae  rufo-piceous,  tarsi  rufous ;  antennae  moder- 
ately long,  joints  three,  four  and  five  rather  elongate,  sixth  triangular  and 
about  as  wide  as  the  fifth 49  fulvipilosus,  n.  sp. 

— .  Antennae,  tibiae  and  tarsi  dark  piceous;  antennae  long,  heavy,  last  two 
joints  particularly  stout;  sixth,  seventh  and  eighth  feebly  incrassate, 
about  equal  in  length  and  size;  maculation  tending  to  form  five  trans- 
verse fasciae  on  elytra,  the  basal  two  of  which  are  usually  broken 

50.  cervicalis,  n.  sp. 

The  following  two  species  have  not  been  recognized  with  certainty 
in  the  material  that  has  passed  through  my  hands.  Both  species  were 
founded  on  females.  The  following  descriptive  outlines  have  been 
epitomized  from  Casey : 

51.  Listrus  plenus   Casey. — Form  elongate-oval,   strongly  convex. 
Color  polished  black,  without  metallic  lustre ;  second  joint  of  the  antennae 
piceous.     Pubescence  short,  sparse,  feebly  persistent  and  with  large  dark 
areas  on  the  elytra. 

Head  sparsely  perforato-punctate,  the  interspaces  flat  and  somewhat 
rugose.  Antennae  slender,  quite  distinctly  longer  than  the  pronotum,  the 
tenth  joint  about  as  long  as  wide. 

Pronotum  two-fifths  wider  than  long ;  sides  evenly  and  moderately 
convergent  and  feebly  arcuate  from  near  the  base  to  the  obtuse  and  some- 
what rounded  apical  angles ;  apex  broadly  arcuato-truncate  and  wider 
than  the  pedunculiform  part  of  the  base,  which  is  more  pronounced  than 
usual ;  disk  widest  at  basal  third,  moderately  coarsely,  deeply  and  very 
closely  perforato-punctate,  scabrous  and  opaque  in  lateral  fourth ;  punc- 
tures separated  by  barely  their  own  dimensions,  the  narrow  interspaces 
sparsely  and  feebly  punctulate.  Elytra  three-fourths  longer  than  wide, 
fully  one-third  wider  than  the  pronotum,  parallel,  acutely  parabolic  in 
apical  third,  coarsely  and  rather  sparsely  punctate,  more  finely  so  toward 
apex  as  usual. 

Length,  3.0  mm. ;  width,  1.2  mm. 

Type  locality. — Vancouver  Island. 

52.  Listrus  densicollis  Casey. — Form  stout,  oblong-oval,  somewhat 
depressed  above.     Color  black  with  a  greenish-brassy  lustre.     Pubescence 
short,  sparse,  cinereous,  not  very  persistent,  with  large  indefinite  dark 
patches  on  the  elytra. 

Head  densely  punctate,  interspaces  feebly  rugose ;  frontal  impressions 
distinct.  Antennae  only  very  little  longer  than  the  pronotum,  tenth  joint 
one-half  wider  than  long. 

Pronotum  one-half  wider  than  long,  widest  at  basal  third,  where  the 
sides  are  rather  broadly  and  evenly  rounded,  thence  strongly  convergent 


SYNOPSIS  OF  SPECIES  OF  LISTRUS  181 

and  feebly  arcuate  to  the  apex,  the  latter  truncate  and  scarcely  narrower 
than  the  pedunculate  base ;  apical  angles  very  obtusely  rounded ;  lateral 
serrules  strong;  disk  somewhat  finely  but  deeply  and  closely  perforato- 
punctate,  the  intervals  flat  and  feebly,  sparsely  punctulate,  only  slightly 
wider  than  the  punctures ;  rugose  area  in  lateral  fourth. 

Elytra  oblong,  two-thirds  longer  than  wide,  fully  one-third  wider  than 
the  pronotum,  parallel,  parabolic  in  apical  third ;  humeri  tumid  and 
prominent;  punctures  rather  coarse,  deep  and  sparse. 

Length,  3.2-3.8  mm. ;  width,  1.2-1 .4  mm. 

Type  locality. — Napa  County,  California. 

In  the  Leng  Catalogue  of  the  Coleoptera  of  North  America  recently 
issued,  Motschulsky's  Listrus  tibialis  has  been  given  a  regular  place  in  the 
arrangement  of  species.  Casey  was  unable  to  identify  it  when  he  wrote 
his  revision  of  the  genus  Listrus  Mots,  in  1895.  I  have  likewise  failed  to 
recognize  it  up  to  the  present  time.  Inasmuch  as  I  have  mentioned  two 
of  Casey's  species  that  I  could  not  identify,  it  will  be  quite  proper  at  the 
present  time  to  give  the  translation  of  Motschulsky's  description,  which 
is  as  follows : 

53.  Listrus  tibialis  Mots. — Form  elongate-subovate,  rather  convex 
and  shining;  punctate  and  sparsely  clothed  with  cinereous  pubescence. 
Color  nigro-aeneous  above,  black  beneath ;  tibiae  rufo-testaceous,  labrum 
and  tarsi  inf uscate ;  antennae  and  femora  blackish. 

Head  between  the  eyes  transversely  carinulate,  carinula  strongly 
shining. 

Pronotum  transverse,  narrowed  anteriorly ;  apical  angles  distinct,  the 
posterior  angles  rounded;  sides  subcrenulate  and  set  with  rather  long 
fimbriae. 

Elytra  subovate  and  arcuately  narrowed  posteriorly;  clothed  with 
cinereous  pubescence. 

Measurements. — Length,  1%  1.;  width,  %  1. 

Habitat. — Ross  (Sonoma  County,  California). 

It  is  hoped  that  the  synoptical  arrangement  as  given  above  will  aid 
the  student  in  placing  the  several  species  of  these  pretty  little  melyrids. 
There  will  be  disappointment  in  its  application,  as  there  are  yet  many 
species  to  be  discovered,  and  these  undescribed  forms  will  unwittingly  be 
intermixed  and  referred  to  some  of  the  species  already  named.  The 
selected  characters  are  not  as  complete  as  I  wish  they  were,  for  a  con- 
siderable per  cent  of  the  specimens  studied  were  mounted  in  such  a  way 
that  it  was  impossible  to  see  the  antennae,  metatrochanters  or  fifth  ventral 
abdominal  segment.  An  attempt  was  made  to  remount  some  of  the 
specimens,  but  they  proved  too  fragile  and  could  not  be  properly  manipu- 
lated without  destruction.  So  it  was  decided  at  the  present  time  not  to 


182      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

delay  making  known  the  species  that  have  already  in  part  been  distributed 
under  manuscript  names. 

Before  closing  this  thesis  it  will  be  well  to  urge  the  student  to 
familiarize  himself  with  the  pronotal  sculpturing,  to  determine  that  it 
consists  of  distinct  and  well  defined  punctures  and  that  the  intervals  may 
be  more  or  less  distinctly  indented  after  the  manner  of  hammered  brass. 
The  basal  joint  of  the  antennae  is  always  black,  with  one  exception,  and 
that  is  in  concurrens  Casey. 

Genus  DASYTES  Payk. 

Dasytes  nevadensis,  new  species. — Form  subcuneiform,  elongate. 
Color  polished  black,  with  a  very  faint  greenish  metallic  lustre ;  tibiae  and 
tarsi  scarcely  or  feebly  nigro-piceous. 

Pubescence  nigro-fuscous,  sparse,  moderate  in  length,  rather  coarse, 
semierect,  very  irregularly  directed  on  the  head  and  pronotum ;  marginal 
cilia  widely  spaced  and  not  noticeably  fimbriform. 

Head  rather  large,  wider  across  the  eyes  than  the  pronotal  apex ;  front 
very  broadly  and  feebly  bi-impressed,  punctures  sparse,  rather  small ; 
surface  shining  and  somewhat  wrinkled  along  the  eyes  and  frontal  margin  ; 
muzzle  short.  Eyes  moderately  prominent,  broadly  convex,  setigerous, 
setae  short  and  sparse.  Antennae  rather  long,  reaching  well  beyond  the 
pronotal  base,  joints  four  to  ten  inclusive  subequal,  eleventh  about  twice 
as  long  as  wide. 

Pronofum  about  a  fourth  wider  than  long,  widest  behind  the  middle 
where  the  sides  are  distinctly  and  evenly  arcuate,  somewhat  constricted 
anteriorly  where  the  sides  are  rather  deeply  sinuate  behind  the  apical 
angles,  lateral  margins  distinctly  and  rather  strongly  serrulate;  apex 
broadly  and  feebly  arcuate,  rather  arcuato-truncate  in  middle  third, 
narrower  than  the  base ;  apical  angles  obtuse  and  more  or  less  rounded ; 
base  broadly  arcuate,  continuously  so  with  the  broadly  rounded  angles, 
which  are  really  obtuse  and  reflexed  when  viewed  obliquely ;  disk  rather 
strongly  convex  anteriorly  and  centrally,  less  so  toward  the  basal  angles, 
submarginal  line  very  strong  and  passing  arcuately  from  side  to  side 
along  the  basal  margin,  central  area  very  sparsely  punctate,  punctures 
quite  evenly  spaced,  intervening  surface  glabrous  and  shining,  lateral  area 
rather  narrow  and  strongly  reto-rugose. 

Elytra  about  one  and  a  half  times  longer  than  wide,  widest  behind 
the  middle,  sides  converging  anteriorly,  apex  rather  broadly  and  para- 
bolically  rounded ;  disk  moderately  convex,  punctures  rather  sparse  and 
not  strong,  somewhat  obscured  by  the  transverse  subrugulation. 

Abdomen  densely  and  rather  coarsely  punctate,  surface  distinctly  and 
transversely  rugulose. 


DASYTES  PAYK,  NEW  SPECIES  OF  183 

Male. — Somewhat  narrow.  Fifth  ventral  segment  truncate,  adjacent 
surface  rather  broadly  and  lunately  impressed ;  pygidium  arcuately  emar- 
ginate,  lateral  angles  rounded. 

Female. — Somewhat  broader.  Fifth  ventral  arcuately  rounded  at 
apex,  sometimes  the  adjacent  surface  is  somewhat  flattened. 

Measurements. — Length,  3.0-3.4  mm.;  width,  1.0-1.4  mm. 

Type  locality. — Goldfield,  Esmeralda  County,  Nevada. 

Holotype,  male,  and  allotype,  female,  in  my  own  collection.  Col- 
lected by  Mr.  F.  W.  Nunenmacher,  who  also  possesses  paratypes. 
Obtained  by  beating. 

Remarks. — By  the  heavy  submarginal  line  and  the  pubescence  not 
being  intermingled  with  longer  erect  hairs  and  black  legs,  it  is  related  to 
'nitens  Casey.  In  nit  ens  however,  the  pronotal  sides  are  much  more 
evenly  arcuate  and  not  constricted  and  emarginate  behind  the  apical 
angles. 

Dasytes  angulatus,  new  species. — Form  elongate  ovate,  wider  pos- 
teriorly, somewhat  depressed.  Color  black,  sometimes  brownish,  moder- 
ately shining ;  mouth  parts  more  or  less  pale,  tibiae  and  tarsi  pale  rufous 
or  rufo-piceous. 

Pubescence  brown  to  grayish-brown,  short,  decumbent  and  sparse ; 
pronotal  and  elytral  fimbriae  very  short  and  somewhat  blackish,  on  the 
former  somewhat  erect,  on  the  latter  a  little  longer  and  paler  at  apex. 

Head  about  as  wide  as  the  pronotal  apex ;  front  feebly  convex,  im- 
pressions feeble,  rather  long  and  separated  by  a  median  and  very  slight 
longitudinal  convexity ;  surface  finely,  sparsely  punctate  and  micro- 
scopically reticulate. 

Pronotum  about  a  fourth  wider  than  long;  sides  scarcely  arcuate, 
rather  broadly  sinuate  behind  the  angles,  convergent  from  base  to  apex 
as  viewed  from  above,  or  feebly  arcuate  at  middle,  straight  and  converging 
anteriorly,  converging  to  base  in  basal  fourth  as  viewed  obliquely  from 
the  side ;  apex  rather  broadly  emarginate ;  apical  angles  blunt  and  sub- 
rectangular ;  base  broadly  arcuate,  distinctly  sinuate  within  the  angles, 
these  are  subacute  and  more  or  less  minutely  prominent ;  disk  with  a  dis- 
tinct submarginal  line  that  is  somewhat  abbreviated  anteriorly  and 
passing  arcuately  into  the  basal  submarginal  groove,  surface  declivous 
laterally,  feebly  impressed  before  and  within  the  basal  angles  so  that  the 
intervening  surface  appears  oblique  and  more  or  less  gable-like  toward 
the  impressed  line  at  basal  fourth ;  central  area  very  feebly  and  sparsely 
punctate,  laterally  microscopically  reticulato-asperate. 

Elytra  about  twice  as  long  as  the  width  at  base;  sides  moderately 
divergent  to  basal  third,  thence  broadly  and  evenly  arcuate  to  apex; 


184      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

margin  narrow,  not  noticeably  reflexed ;  disk  very  gradually  declivous 
posteriorly,  more  strongly  so  laterally,  surface  very  finely  and  reticulato- 
rugulose;  punctures  fine  and  moderately  sparse;  humeri  not  prominent 
and  without  umbones. 

Abdomen  very  finely  punctate. 

Male. — Fifth  ventral  segment  rather  broadly  rounded  but  truncate  at 
middle  third  of  apex;  surface  rather  broadly  impressed  before  the  trun- 
cature,  margin  set  with  short  stiff  hairs. 

Female. — Broader.  Pronotal  sides  more  broadly  arcuate  and  less 
convergent  toward  apex,  surface  not  noticeably  impressed  in  the  basal 
region  and  more  evenly  convex,  submarginal  line  less  deeply  impressed 
toward  base.  Fifth  ventral  broadly  rounded. 

Measurements. — Length  (Types),  2.8-3.1  mm.;  width,  1.2-1.4  mm. 

Holotype,  male,  and  allotype,  female,  in  the  collection  of  the  Cali- 
fornia Academy  of  Sciences.  Paratypes  in  the  Academy's  and  my  own 
collection. 

Type  locality. — Mt.  Eddy,  Siskiyou  County,  California.  Collected  by 
Mr.  E.  P.  Van  Duzee,  on  July  28,  1918,  at  an  elevation  of  9000  feet. 

In  many  specimens  the  trochanters  and  anterior  femora  are  pale 
dorsally  at  base.  The  first  joint  of  the  hind  tarsi  is  about  equal  in  length 
to  the  third  and  fourth  taken  together.  Angulatus  by  its  distinct  basal 
angles  of  the  pronotum  differs  from  all  other  species  of  the  genus,  its 
pubescence  is  uniform  over  the  upper  surface,  longer,  denser  and  grayer 
on  the  abdomen  beneath. 

Dasytes  shastensis  horridulus,  new  subspecies. — Size  comparatively 
large  and  quite  similar  in  the  sexes.  Form  oblong-oval,  about  two  and  a 
seventh  times  longer  than  wide,  moderately  convex.  Color  black ;  mouth- 
parts,  antennae,  tibiae  and  tarsi  more  or  less  piceous  ;  surface  more  or  less 
shining,  metallic  lustre  scarcely  discernible. 

Pubescence  quite  long  and  abundant,  consisting  of  erect,  black, 
bristling  and  paler  almost  recumbent  hairs. 

Head  scarcely  as  wide  as  the  pronotal  apex ;  front  quite  plane,  feebly 
and  broadly  impressed ;  quite  coarsely  and  closely  punctate.  Muzzle  short 
and  broad.  Eyes  large  and  moderately  convex,  distinctly  setigerous. 

Antennae  similar  in  the  sexes,  rather  long  and  loose-jointed,  rather 
moderately  compressed,  not  incrassate ;  second  and  third  joints  subequal 
in  length,  second  about  a  half  longer  than  wide,  nearly  twice  as  stout  as 
the  third,  which  is  relatively  slender  and  almost  twice  as  long  as  wide ; 
fourth  and  fifth  joints  subequal  in  length,  subtriangular  and  slightly  longer 
than  wide ;  sixth  to  the  tenth  inclusive  triangulo-moniliform,  about  as 
long  as  wide;  eleventh  elongate,  subovate  and  distinctly  as  long  as  wide. 


SYNOPSIS  OF  SPECIES  OF  DASYTES  185 

Pronotum  about  a  fourth  wider  than  long ;  apex  feebly  arcuate ;  apical 
angles  quite  broadly  and  obtusely  rounded ;  sides  evenly  and  moderately 
strongly  arcuate,  serrations  short  and  blunt ;  base  broadly  and  rather 
strongly  arcuate,  rounding  into  the  sides  without  basal  angles ;  disk 
evenly  and  moderately  strongly  arcuate  from  side  to  side,  punctures 
moderate  in  size,  evenly  placed,  separated  by  a  distance  equal  to  their 
diameter,  submarginal  line  coarse,  sometimes  more  or  kss  broken  ante- 
riorly, lateral  area  reticulato-rugose. 

Elytra  oblong,  a  little  wider  posteriorly,  sides  parallel,  very  feebly 
arcuate,  apex  very  broadly  rounded ;  punctures  distinctly  smaller  than  on 
the  pronotum,  and  separated  by  a  distance  equal  to  twice  their  diameter ; 
base  equal  to  the  width  of  the  pronotum. 

Abdomen  finely  and  rather  densely  punctate;  fifth  segment  more 
densely  and  subasperately  sculptured. 

Legs  moderate  in  length  and  not  stout ;  metatarsi  long  and  slender, 
quite  equal  to  the  length  of  their  tibia. 

Male. — Fifth  ventral  segment  broadly  and  feebly  arcuato-truncate. 

Female. — Fifth  ventral  broadly  rounded  at  apex. 

Measurements. — Length   (Types),  3.8-4  mm.;  width,   1.7-1.8  mm. 

Holotype,  male,  and  allotype,  female,  and  one  paratype  (male)  in  my 
own  collection. 

Type  locality. — Crystal  Lake,  San  Mateo  County,  California.  Col- 
lected by  myself  on  June  27,  1919. 

Distribution. — Occurs  also  on  Mt.  Tamalpais,  Marin  County,  Cali- 
fornia ;  captured  June  25,  1920. 

Quite  distinct  in  its  large  size,  longer  and  moderately  close 
pubescence,  stouter  and  more  elongate  second  antennal  joint ;  the  sub- 
marginal  pronotal  line  is  coarse  and  oftentimes  much  broken  up  anteriorly. 
It  is  undoubtedly  closely  related  to  shastensis  Blais.,  which  when  com- 
pared with  horridulus,  is  smaller  in  size,  pronotum  less  strongly  punctured 
in  the  central  area,  second  antennal  joint  less  elongate,  less  stout  and 
inclined  to  narrow  toward  the  base. 

SYNOPSIS  OF  THE  SPECIES  OF  DASYTES  PAYK. 

Submarginal  line  of  the  pronotum  feeble,  frequently 'only  evident  as  an  abrupt 

division  between  the  smooth  and  rugose  portions  of  the  surface 1 

Submarginal  line  strong,  impressed  and  usually  entire 2 

1.  Elytra  each  with  longitudinal  dark  vittiform  lines 1.     lineellus  Casey 

— .     Elytra   without  vittae 1 1 

2.  Pubescence  distinctly  intermingled  with  longer,  erect  black  hairs 3 

— .     Pubescence  not  intermingled  with  longer,  erect  black  hairs 8 

3.  Elytral  vestiture  blackish  throughout 2.    hudsonicus  Lee. 


186      NEW  SPECIES   MELYRIDAE.  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

— .     Elytral  vestiture  more  or  less  pale 4 

4.  Vestiture  cinereous  but  becoming  dark  and  inconspicuous  behind  the  middle 

of  the  elytra  which  are  broad  and  depressed 3.     expansus  Casey 

— .     Vestiture  cinereous  with  longer,  erect  black  hairs  on  the  elytra 5 

5.  Basal  angles  broadly  rounded;  pale  hairs  recumbent 6 

— .     Basal  angles  obtuse  7 

6.  Size  usually  under  3  mm. ;  pubescence  shorter 4.     shastensis  Blais. 

— .    Size  larger,  4  mm. ;  pubescence  longer,  form  more  oblong 

5.     horridulus,  n.  subsp. 

7.  Thoracic  punctures  coarse  and  impressed ;  pale  hairs  suberect..  6.    vicinus  Blais. 
— .    Thoracic  punctures  fine  and  sparse 7.    obtusus  Casey 

8.  Legs  pale  throughout 9 

— .    Legs  bicolored,   femora  black 10 

9.  Pronotal  margins  serrulate  and  deeply  sinuate  behind  the  apical  angles, 

pubescence  blackish  8.    nevadensis,  n.  sp. 

— .     Pronotal  margins  feebly  serrulate  and  feebly  sinuate  anteriorly ;  pubescence 

yellowish  9.    nitens  Casey 

10.  Elytral  pubescence  blackish  and  inconspicuous  but  becoming  ashy  toward 

base;  antennae  stouter  and  longer 10.     breviusculus  Casey 

— .     Elytral    pubescence    pale   cinereous    throughout,    even    but   rather   sparse ; 

body  more  elongate;  antennae  shorter 11.     dissimilis  Casey 

11.  Elytral  vestiture  uneven  in   distribution,  brownish-black  hairs   forming  a 

basal  and  subapical  broad  fascia _ 12.    seminudus  Lee. 

— .  Elytral  vestiture  uniform   in  distribution 12 

12.  Basal  angles  of  pronotum  rounded 13 

— .  Basal  angles  more  or  less  distinct 17 

13.  Legs  black  or  blackish  in  color  throughout 14 

— .  Legs  more  or  less  pale  in  color 15 

14.  Pubescence  coarse,  cinereous,  dense,  short,  recumbent,  and  uniformly  dis- 

tributed   13.    musculus  Fall 

— .  Pubescence  cinereous,  moderately  dense,  rather  short,  not  very  coarse, 
suberect  and  intermixed  toward  sides  with  a  few  black  setae  on  pro- 
notum   14.  macer  Casey 

15.  Legs  bicolored,  pale  rufous,  femora  black 15.    cruralis  Lee. 

— .    Legs  rufo-ferruginous  throughout,  femora  occasionally  slightly  darker 16 

16.  Prothorax  much  narrower  than  the  base  of  the  elytra... „ 16.    pusillus  Lee. 

— .     Prothorax  equal  in  width  to  base  of  elytra 17.     minutus  Casey 

17.  Basal  angles  of  pronotum  subrectangular  and  distinct,  base  sinuate  within 

the  angles ;  legs  pale 18.     angulatus  n.  sp. 

— .     Basal  angles  obtuse  18 

18.  Legs  dark  in  color;  elytral  vestiture  consisting  of  shorter  and  longer  hairs 

which  are  confusedly  intermingled '. 19.    fastidiosus  Casey 

— .     Legs  entirely  pale  or  bicolored 19 

19.  Legs  bicolored,  femora  black ;  erect  hairs  of  elytra  extremely  short  and  in- 

distinct, only  visible  toward  apex 20.     depressulus  Casey 

— .  Legs  rufo-testaceous ;  pubescence  luteo-cinereous,  rather  long,  suberect, 
consisting  especially  on  elytra  of  somewhat  uniformly  mixed  shorter, 

more  inclined  and  recurved,  longer,  and  a  little  less  inclined  hairs 

21.     dementae   Fall 


NOTES  ON   DISTRIBUTION  OF  SPECIES  OF  DASYTES  187 

NOTES  ON  THE  DISTRIBUTION  OF  THE  SPECIES  OF  DASYTES  PAYK. 

1.  Dasytes  lineellus  Casey. — California:     Los  Angeles  County   (Casey);   Mount 

Wilson,  June  18th,  (Fall)  ;  Bear  Lake,  San  Bernardino  County,  June  5,  1919, 
(J.  O.  Martin)  ;  Hesperia,  June  30,  1918,  (J.  O.  Martin)  ;  Mokelumne  Hill, 
Calaveras  County,  July,  elevation  2300  feet,  (Blaisdell)  ;  Mariposa  County, 
June  17,  1914,  (F.  W.  Nunenmacher). 

2.  Dasytes    hudsonicus    Lee. — Hudson     Bay    Territory.     Colorado :       Leadville, 

July  7-14,  1896,  elevation  10,000-11,000  feet,  (Wickham)  ;  Clear  Creek, 
June  2,  1881,  (Liebeck).  Arizona  (Casey). 

3.  Dasytes  expansus  Casey. — California:     North  of  San  Francisco  (Casey). 

4.  Dasytes  shastensis  Blais. — California:     Shasta  Retreat,  SSskiyou  County,  ele- 

vation 2416  feet,  July,  1905,  (Blaisdell)  ;  Clayton,  Shasta  County,  July  13, 
1918,  (E.  P.  Van  Duzee)  ;  Fairfax,  Marin  County,  June,  (Blaisdell)  ;  Mari- 
posa County,  June.  • 

5.  Dasytes    shastensis    horridulus,    new    subspecies. — California :      Crystal    Lake, 

San  Mateo  County,  June  27,  1919,  (Blaisdell)  ;  Mount  Tamalpais,  Marin 
County,  June  25,  1919,  (Blaisdell). 

6.  Dasytes  vicinus  Blais. — California.     San  Diego,  (Blaisdell). 

7.  Dasytes  obtusus  Casey. — Colorado:     Glenwood  Springs,  July,  (Dr.  A.  Fenyes). 

8.  Dasytes    nevadensis,    new    species. — Nevada :     Goldfield,    Esmeralda    County, 

June  6,  1908,  (F.  W.  Nunenmacher). 

9.  Dasytes  nitens  Casey. — California:     Duncan  Mills,  Sonoma  County,  June  30, 

1908,  (Blaisdell)  ;  Samoa,  Humboldt  County,  June  21,  1916,  (Blaisdell)  ; 
Marin  County,  (Casey). 

10.  Dasytes   breviusculus   Casey. — California:     Sisson,   Siskiyou  County,   July  26, 

1918,  (E.  P.  Van  Duzee)  ;  Clayton,  July  17,  1918,  (Van  Duzee). 

11.  Dasytes  dissimilis  Casey. — California:     San  Bernardino  Mountains,  (Fall). 

12.  Dasytes  musculus  Fall.— Calif ornia :     Mount  Wilson,  July  23,  1905 ;  Pasadena, 

Pomona  and  Riverside,  (Fall). 

13.  Dasytes  seminudus  Lee. — California :     Vine  Hill,  Contra  Costa  County,  June  7, 

1908,  (Blaisdell)  ;  Davis  Meadow  near  Railroad  Flat,  Calaveras  County, 
July,  1918,  elevation  2800  feet,  (Blaisdell)  ;  Mount  Diablo,  July  16th, 
W.  M.  Giffard)  ;  Mojave,  in  May. 

14.  Dasytes  macer  Casey. — Southern  California  (Casey). 

15.  Dasytes  cruralis  Lee.— Calif  ornia :     Mokelumne  Hill,  Calaveras  County,  June, 

1898,  (Blaisdell)  ;  Shasta  Retreat,  Siskiyou  County,  July,  elevation  2416  feet, 
(Blaisdell)  ;  Yosemite,  Mariposa  County,  June;  Mariposa  County,  June  16th, 
W.  M.  Giffard)  ;  Tuolumne  County,  May  14,  1914,  (F,  W.  Nunenmacher)  ; 
Duncan  Mills,  Sonoma  County,  July  14,  1908,  (Blaisdell)  ;  Lodi  and  Stockton, 
San  Joaquin  County,  May  14,  1911,  (Blaisdell).  Oregon:  Colestin,  Jackson 
County,  July  31,  1918,  (E.  P.  Van  Duzee). 

16.  Dasytes  pusillus  Lee. — California :     San  Diego,    (Casey)  ;   Palm  Springs,  Im- 

perial County,  May,  (Dr.  Fenyes)  ;  Mecca,  Colorado  Desert,  April  13,  1917, 
(J.  O.  Martin). 

17.  Dasytes  minutus  Casey. — California.     (Casey). 

18.  Dasytes  angulatus,  new  species. — California:     Mount  Eddy,  9000  feet,  July  28, 

1918,  (E.  P.  Van  Duzee). 

19.  Dasytes  fastidiosus   Casey.— California :      Sausalito,   Marin   County,  April  26, 

1914,  (Blaisdell)  ;  Vine  Hill  near  Martinez,  Contra  Costa  County,  June  7, 
1908,'  (Blaisdell). 


188      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

20.  Dasytes  depressulus  Casey. — Nevada:     (Casey). 

21.  Dasytes  dementae  Fall. — Island  of  San  Clemente:     (Fall). 

LISTRIMORPHA,  New  Genus. 

Form  of  a  female  Eschatocrepis  constrictus  Lee.,  sparsely  and  uni- 
formly pubescent.  Pronotum  without  a  submarginal  line,  lateral  margins 
finely  serrulate  and  feebly  fimbriate ;  disk  broadly  impressed  within  the 
basal  angles.  Last  joint  of  the  maxillary  palpi  conical. 

Antennae  with  a  feeble  three-jointed  club ;  fifth  joint  larger  than  the 
fourth  and  angulate  anteriorly  nearly  as  in  Listrus;  joints  sixth,  seventh 
and  eighth  much  smaller  than  the  fifth  and  narrower  than  those  of  the 
club. 

Epipleura  rather  narrow  at  base,  gradually  narrowing  to  become 
evanescent  behind  the  middle,  superior  margin  feeble. 

Ungual  appendages  slightly  longer  than  the  length  of  the  claws, 
attached  nearly  throughout  their  length;  equal,  subacute  on  the  middle 
and  posterior  claws. 

Legs  slender.  Claws  slender.  Tarsi  slender,  third  and  fourth  joints 
together  about  equal  to  the  fifth,  the  latter  and  the  first  subequal ;  second, 
two-thirds  as  long  as  the  first.  Femora  not  stout,  somewhat  compressed. 

Listrimorpha  pallipes,  new  species. — Form  elongate  ovate,  wider  pos- 
teriorly. Color  black,  somewhat  dull  to  shining.  Antennae,  except  the 
first  joint  which  is  dark,  and  legs  testaceous;  labrum  yellowish,  femora 
more  or  less  piceous  toward  base.  Surface  microscopically  reticulato- 
rugulose.  Pubescence  short,  sparse,  recumbent  and  pale  in  color,  not 
conspicuous;  elytral  margins  loosely  fimbriate,  hairs  longer  on  the  apex. 

Head  moderate,  as  wide  as  the  pronotal  apex ;  front  feebly  convex, 
feebly  bi-impressed,  impressions  somewhat  elongate,  beginning  on  a  line 
opposite  the  antennal  base ;  vertex  obsoletely  impressed  at  the  middle ; 
very  finely  and  sparsely  punctate.  Eyes  rather  large  and  somewhat 
strongly  convex. 

Antennae  reaching  to  about  the  pronotal  base ;  first  joint  smaller  than 
in  Listrus,  second  evenly  oval  and  about  equal  in  diameter  to  the  first ; 
third  and  fourth  somewhat  compressed ;  third  obconical  a  little  shorter 
than  the  second;  fourth  subtriangular  (isosceles)  and  angulate  anteriorly; 
fifth  larger  and  similarly  angulate  anteriorly,  as  long  as  the  second  and 
distinctly  larger  than  either  the  sixth,  seventh  or  eighth;  sixth  smallest 
and  about  as  long  as  wide,  two-thirds  as  wide  as  the  fifth,  seventh  a  little 
larger  and  triangulo-oval ;  eighth  about  as  long  as  the  sixth  and  slightly 
wider,  sides  more  parallel  and  more  arcuate,  about  as  long  as  wide; 
eleventh  pointed  oval  and  a  half  longer  than  wide. 


LISTRIMORPHA,  NEW  GENUS  189 

Pronotum  transverse,  about  a  third  wider  than  long,  base  slightly 
wider  than  the  apex;  sides  angulate  just  slightly  in  front  of  the  middle, 
thence  quite  straight  and  convergent  to  apex,  straight,  convergent  or 
parallel  posteriorly  to  base,  .margin  serrulate;  apex  feebly  emarginate; 
apical  angles  obtuse  and  very  slightly  rounded;  basal  angles  distinct, 
obtuse  to  rectangular  and  somewhat  reflexed;  base  transverse  in  middle 
two-fourths,  thence  oblique  and  feebly  sinuate  to  the  basal  angles ;  disk 
moderately  convex,  rather  evenly  so  in  central  area  and  at  apex,  more  or 
less  broadly  and  distinctly  impressed  in  the  region  of  the  basal  angles  and 
along  the  sides  to  the  angulation,  and  very  feebly  so  across  in  front  of  the 
base,  slightly  constricted  laterally  behind  the  apex,  very  finely  and 
sparsely  punctate,  slightly  asperate  in  the  lateral  area. 

Elytra  oblong,  slightly  dilated  behind  the  middle,  about  twice  as  long 
as  wide,  feebly  convex  on  the  disk,  more  strongly  so  laterally;  apices 
slightly  dehiscent,  angles  obtuse ;  finely  and  sparsely  punctate.  Scutellum 
transversely  oblong. 

Under  surface  of  the  body  finely  sculptured. 

Measurements. — Length,  2.6  mm. ;  width,  1.0  mm. 

Holotype,  male,  and  allotype,  female,  in  my  own  collection ;  paratypes 
in  that  of  Mr.  F.  W.  Nunenmacher. 

Type  locality. — Goldfield,  Esmeralda  County,  Nevada.  Collected  by 
Mr.  Nunenmacher  on  June  29,  1907.  Five  specimens  studied. 

Remarks. — This  species  is  distinct  in  having  the  pronotum  without  a 
submarginal  line  or  lateral  rugose  area  and  short  ungual  appendages. 
In  Casey's  table  of  the  Melyrid  tribe  Dasytini  it  falls  between  Dasytastes 
and  Eschatocrepis,  and  by  its  general  facies  and  structural  characters  it  is 
more  closely  related  to  the  latter  genus.  From  both  Listrus  and  Eschato- 
crepis it  can  be  recognized  by  the  subangulate  sides  of  the  pronotum,  the 
latter  less  convex  and  with  serrulate  lateral  margins. 

The  fifth  ventral  segment  has  the  apical  margin  set  with  a  line  of 
short  and  rather  stiff  hairs.  Sexual  differences  are  not  very  evident  in 
the  small  series  at  hand. 

Listrimorpha  should  precede  Eschatocrepis  in  our  lists. 

Dasytastes  vanduzeei,  new  species. — Form  subovate.  Color  black; 
head,  prothorax,  legs  and  basal  portion  of  the  antennae,  rufo-f ulvous ; 
metasternum  more  or  less  rufous. 

Pubescence  rather  short  and  recumbent,  quite  dense  on  the  elytra, 
somewhat  less  so  on  the  pronotum  and  under  surface  of  the  body. 
Abdominal  surface  dull.  Elytra  feebly  aeneous  and  maculate  with  several 
slightly  elongate  blackish  spots,  those  of  the  apical  half  anastomosing  to  a 
greater  or  less  extent. 


190      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Front  of  the  head  flat,  finely  and  evenly  punctate,  eyes  not  prominent. 

Pronotum  finely  and  evenly  punctate ;  apex  arcuato-truncate ;  base 
arcuate ;  sides  more  strongly  arcuate  in  the  basal  half,  thence  broadly  and 
evenly  so  to  apex,  very  slightly  sinuate  just  before  the  almost  rounded 
basal  angles,  serrulations  small ;  apical  angles  rounded ;  margins  fimbriate, 
fimbriae  not  long. 

Elytra  finely  punctate,  apical  margins  serrulate. 

Measurements. — Length,  2.2  mm. ;  width,  1.0  mm. 

Type  locality. — Palm  Springs,  Riverside  County,  California.  Col- 
lected on  May  20th,  by  Mr.  E.  P.  Van  Duzee. 

Described  from  two  females,  both  in  the  collection  of  the  California 
Academy  of  Sciences. 

Remarks. — Vanduzeei  differs  from  bicolor  Casey  in  the  color  and 
maculation  of  the  elytra.  In  bicolor  the  elytral  pubescence  is  blackish, 
while  in  vanduzeei  it  is  pale,  except  in  spots  as  stated  above.  Vanduzeei 
should  follow  bicolor  in  our  lists. 

BIBLIOGRAPHY  AND  CABINET  ARRANGEMENT. 

1.  Listrus  definitus  Fall. 

"Occasional  Papers,"  Calif.  Acad.  Sci's,  8,  248,  1901. 

2.  Listrus  cephalicus,  new  species. 

3.  Listrus  senilis  Lee. 

Dasytes  senilis  Lee.,  Proc.  Acad.  Nat.  Sci's,  Phila.,  6,  170. 
Proc.  Acad.  Nat.  Sci's,  Phila.,  358,  Dec.,  1866. 

Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  551, 
July,  1895. 

4.  Listrus  uniformis  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  ScL  8,  552,  July,  1895. 

5.  Listrus  davicornis  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  552,  July,  1895. 

6.  Listrus  coloradensis,  new  species. 

7.  Listrus  obscurellus  Lee. 

Dasytes  obscurellus  Lee.,  Proc.  Acad.  Nat.  Sci's,  Phila.,  6,  170. 

Proc.  Acad.  Nat.  Sci's,  Phila.,  358,  Dec.,  1866. 

Casey,  Thos.  L.— XV.— "Coleopt.  Not,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  553, 

July,  1895. 

'8.     Listrus  punctatus  Motsch. 
Bull.  Mosc.,  1859. 
XV.— Casey,  Thos.  L.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  543, 

July,  1895. 
9.    Listrus  dilutus,  new  species. 

10.  Listrus  martini,  new  species. 

11.  Listrus  annulatus  Casey. 

XV.— "Coleopt.  Not,  VI.,"  Annals  N.  Y.  Acad.  Sci.,  8,  559,  July,  1895. 

12.  Listrus  rubripes  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  560,  July,  1895. 


BIBLIOGRAPHY    AND    CABINET    ARRANGEMENT  191 

13.  Listrus  maculosus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  550,  July,  1895. 

14.  Listrus  montanus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  546,  July,  1895. 

15.  Listrus  confusus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  557,  July,  1895. 

16.  Listrus  extricatus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  554,  July,  1895. 

17.  Listrus  difficilis  Lee. 

Dasytes  difficilis  Lee.,  Proc.  Acad.  Nat.  Sci's,  Phila.,  6,  170.    Proc.  Acad.  Nat. 

Scis,  Phila.,  358,  Dec.,  1866. 
Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  555, 

July,  1895. 

18.  Listrus  fidelis  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  556,  July,  1895. 

19.  Listrus  subaeneous  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  549,  July,  1895. 

20.  Listrus  vestitus,  new  species. 

21.  Listrus  incertus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  559,  July,  1895. 

22.  Listrus  bifasciatus,  new  species. 

23.  Listrus  pardalis  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  551,  July,  1895. 

24.  Listrus  motschulskii  Lee. 

Proc.  Acad.  Nat.  Sci's,  Phila.,  357,  Dec.,  1866. 
Dasytes  canescens  Lee.,  Proc.  Acad.  Nat.  Sci's,  Phila.,  6,  170. 
Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  545, 
July,  1895. 

25.  Listrus  simplex,  new  species. 

26.  Listrus  inters titialis  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  545,  July,  1895. 

27.  Listrus  variegatus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  558,  July,  1895. 

28.  Listrus  incestus,  new  species. 

29.  Listrus  tritus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  557,  July,  1895. 

30.  Listrus  giffardi,  new  species. 

31.  Listrus  parvicollis,  new  species. 

32.  Listrus  luteipes  Lee. 

Dasytes  luteipes  Lee.,  Proc.  Acad.  Nat.  Sci's,  Phila.,  6,  170.    Proc.  Acad.  Nat. 

Sci's,  Phila.,  358,  Dec.,  1866. 
Casey,  Thos.  L.^XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  561, 

July,  1895. 

33.  Listrus  famelicus  Casey. 

.     XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  563,  July,  1895. 

34.  Listrus  ornatulus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  563,  July,  1895. 

35.  Listrus  concurrens  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  561,  July,  1895. 


192      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

36.  Listrus  balteellus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  562,  July,  1895. 

37.  Listrus  tro  chant  erica,  new  species. 

38.  Listrus  liebecki,  new  species. 

39.  Listrus  olympianus,  new  species. 

40.  Listrus  occidens,  new  species. 

41.  Listrus  angulatus,  new  species. 

42.  Listrus  salicis,  new  species. 

43.  Listrus  niveicanthus,  new  species. 

tincticornis,  new  subspecies. 

44.  Listrus  amplicollis  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  547,  July,  1895. 

45.  Listrus  interruptus  Lee. 

Proc.  Acad.  Nat.  Sci.,  Phila.,  357,  Dec.,  1866. 

Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  547, 
July,  1895. 

46.  Listrus  elegantulus,  new  species. 

47.  Listrus  rotundicollis  Casey. 

Dasytes  rotundicollis  Lee.,   Proc.   Acad.  Nat.  Set's,    Phila.,   6,   170.    Proc. 

Acad.  Nat.  Sci's,  Phila.,  358,  Dec.,  1866. 
Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  544, 

July,  1895. 

48.  Listrus  canescens  Mann. 

Dasytes  canescens  Mann.,  Bull.  Mosc.,  247,  1843.    Motsch. :  1.  c.,  2,  391,  1859. 

Le  Conte.— Proc.  Acad.  Nat.  Sci's,  Phila.,  358,  Dec.,  1866. 
Casey,  Thos.  L.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  555,  July, 

1895. 

49.  Listrus  fulvipilosus,  new  species. 

50.  Listrus  cervicalis,  new  species. 

51.  Listrus  plenus  Casey. 

XV.— "Coleopt.  Not,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  549,  July,  1895. 

52.  Listrus  densicollis  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  548,  July,  1895. 

53.  Listrus  tibialis  Motsch. 

Bull.  Mosc.,  2,  391,  1859.    Casey,  Thos.  L.— "Coleopt.  Not.,  VI,"  Annals  N.  Y. 
Acad.  Sci.,  8,  543,  July,  1895. 


1.  Dasytes  lineellus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  B,  581,  Aug.,  1895. 

2.  Dasytes  hudsonicus  Lee. 

Proc.  Acad.  Nat.  Sci.,  Phila.,  360,  1866. 

Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  572, 
Aug.,  1895. 

3.  Dasytes  expansus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  574,  Aug.,  1895. 

4.  Dasytes  shastensis  Blais. 

Ent.  News,  75,  March,  1906. 

5.  Dasytes  shastensis  horridulus,  new  variety. 


BIBLIOGRAPHY    AND    CABINET    ARRANGEMENT  193 

6.  Dasytes  vicinus  Blais. 

Ent.  News,  74,  March,  1906. 

7.  Dasytes  obtusus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  573,  Aug.,  1895. 

8.  Dasytes  nevadensis,  new  species. 

9.  Dasytes  nitens  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  574,  Aug.,  1895. 

10.  Dasytes  breviusculus  Motsch. 

Bull.  Mosc.,  2,  396,  1859. 

Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  576, 
Aug.,  1895. 

11.  Dasytes  dissimilis  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  576,  Aug.,  1895. 

12.  Dasytes  seminudus  Lee. 

Proc.  Acad.  Nat.  Sci.,  Phila.,  360,  1866. 

Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Set.,  8,  581, 
Aug.  1895. 

13.  Dasytes  musculus  Fall. 

"Occasional  Papers,"  Calif.  Acad.  Sci's,  8,  249,  1901. 

14.  Dasytes  tnacer  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  577,  Aug.,  1895. 

15.  Dasytes  cruralis  Lee. 

Pristoscelis  cruralis  Lee.,  Proc.  Acad.  Nat.  Sci's.,  Phila,  355,  1866. 
Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  579, 
Aug.,  1895. 

16.  Dasytes  pusillus  Lee. 

Proc.  Acad.  Nat.  Sci's.,  Phila.,  6,  170;  1.  c.,  360,  1866. 

Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  579, 
Aug.  1895. 

17.  Dasytes  minutus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  580,  Aug.,  1895. 

18.  Dasytes  angulatus,  new  species. 

19.  Dasytes  fastidiosus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  577,  Aug.,  1895. 

20.  Dasytes  depressulus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  579,  Aug.,  1895. 

21.  Dasytes  clementae  Fall. 

"Occasional  Papers,"  Calif.  Acad.  Sci's,  8,  250,  1901. 


Listrimorpha,  new  genus. 
1.    Listrimorpha  pallipes,  new  species. 


1.  Dasytastes  catalinae  Lee. 

Proc.  Acad.  Nat.  Sci's,  Phila.,  361,  1866. 

Casey,  Thos.  L. — XV. — "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  583, 
Aug.,  1895. 

2.  Dasytastes  remissus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  583,  Aug.,  1895. 


194      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

3.  Dasytastes  otiosus  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  584,  Aug.,  1895. 

4.  Dasytastes  dispar  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  585,  Aug.,  1895. 

5.  Dasytastes  ruficollis  Ulke. 

Dasytes  ruficollis  Ulke,  Geog.  and  Geol.  Exp.  and  Surv.,  4to.,  Wheeler,  5,  812, 

1875. 
Casey,  Thos.  L.— XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  586, 

Aug.,  1895. 

6.  Dasytastes  bicolor  Casey. 

XV.— "Coleopt.  Not.,  VI,"  Annals  N.  Y.  Acad.  Sci.,  8,  586,  Aug.,  1895. 

7.  Dasytastes  vanduzeei,  new  species. 

8.  Dasytastes  insularis  Fall. 

"Occasional  Papers,"  Calif.  Acad.  Sci's,  8,  251,  1901. 


CHRYSOMELIDAE  195 


PART  II. 
CHRYSOMELIDAE. 

Glyptoscelis  sequoiae,  new  species. — Form  oblong-oval,  broadly 
rounded  posteriorly,  moderately  convex ;  head  and  prothorax  narrowing 
anteriorly.  Cdlor  a  beautiful  cupreous ;  antennae  more  or  less  rufous, 
terminal  three  joints  black,  seventh  and  eighth  blackish  in  about  apical 
half ;  legs  rufous.  Surface  more  or  less  shining,  sparsely  and  quite  evenly 
clothed  with  a  rather  long,  soft  fulvous  pubescence  which  is  recumbent, 
subsquamiform  and  linear,  each  hair  marked  with  a  longitudinal  median 
stria. 

Head  and  pronotum  somewhat  coarsely,  regularly  punctate ;  punc- 
tures well  denned,  separated  by  a  distance  equal  to  one-half  their 
diameter.  Head  with  an  impunctate  median  line,  front  broadly,  trans- 
versely but  not  strongly  prominent  at  the  clypeo-frontal  junction,  broadly 
and  more  or  less  moderately  impressed  between  the  eyes.  Antennae  long 
and  slender. 

Pronotum  a  little  wider  than  long;  sides  rather  evenly  arcuate, 
moderately  convergent  anteriorly  as  viewed  from  above ;  apex  quite 
broadly  arcuate ;  base  slightly  lobed  at  middle,,  feebly  sinuate  laterally  and 
scarcely  wider  than  the  disk  at  middle;  pubescence  slightly  denser  laterally 
and  more  hair-like  centrally. 

Elytra  oblong,  about  one-half  longer  than  wide,  sides  parallel,  scarcely 
arcuate,  and  broadly,  evenly  arcuate  in  apical  one-fourth ;  disk  evidently 
flattened  about  the  scutellum;  pubescence  appearing  feebly  vittate  when 
viewed  longitudinally ;  punctures  evenly  and  rather  closely  placed,  some- 
what smaller  than  on  the  pronotum,  finer  toward  apex. 

Pubescence  of  under  surface  somewhat  whitish,  scarcely  longer  than 
on  the  supper  surface,  hair-like  and  apparently  not  striate.  Punctuation 
finer  than  above. 

Claivs  cleft.  Inner  division  of  each  much  shorter  and  smaller  than 
the  outer  part,  acute,  especially  on  the  middle  and  posterior  claws. 

Male. — Fifth  ventral  abdominal  segment  somewhat  arcuately 
deflexed  apically,  and  with  a  somewhat  broad,  oblong-oval  impression  at 
middle,  which  may  be  more  or  less  impunctate  at  bottom. 

Female. — Fifth  ventral  distinctly  arcuately  deflexed  apically,  and  more 
or  less  concave  throughout  the  width. 

Measurements. — Length  (Types),  6-7  mm. ;  width,  3-3.5  mm. 


196      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Holotype,  male,  and  allotype,  female,  in  the  collection  of  the  Cali- 
fornia Academy  of  Sciences.  Paratypes  in  that  of  the  Academy's  and  my 
own  collection. 

Type  locality. — Cazadero,  Sonoma  County,  California.  Collected  on 
April  12,  1918,  by  Mr.  E.  P.  Van  Duzee. 

Collected  from  the  foliage  of  Sequoia  sempervirens  Endl.  It  appears 
somewhat  probable  that  when  the  species  of  Glyptoscelis  are  better  known, 
the  food  plant  will  be  found  to  be  more  or  less  different  in  case  of  each 
species.  Glyptoscelis  illustris  Cr.  is  usually  taken  from  yellow  pine  (Pinus 
ponderosa  Dougl.),  in  the  vicinity  of  Mokelumne  Hill,  Calaveras  County, 
California ;  squamulatus  Cr.  is  found  plentifully  on  willow  and  a  closely 
related  species  is  found  on  a  composite  shrub  growing  along  the  San 
Diego  River  in  Mission  Valley,  San  Diego  County,  California.  Two 
species  were  taken  from  juniper  trees  on  Black  Mountain  (elevation  8100 
feet),  at  Huntington  Lake,  Fresno  County,  California,  during  the  month 
of  July,  1919;  another  from  fir  in  Humboldt  County,  California;  finally. 
Mr.  Van  Duzee  has  taken  the  species  described  below  from  wild  rose 
near  Sacramento,  California.  Other  phases  are  at  hand  the  food  plants 
of  which  are  not  known.  Ulke  has  reported  that  the  eastern  species 
pubescens  Fab.  and  barbatus  Say  are  common  on  pine. 

Glyptoscelis  parvulus,  new  species. — Form  rather  short  oblong-oval, 
robust  and  moderately  convex ;  prothorax  distinctly  narrower  than  the 
elytra.  Color  black,  with  an  aeneous,  cupreous  or  virido-aeneous  tinge ; 
legs  rufo-piceous,  palpi  and  antennae  more  or  less  rufous.  Surface  more 
or  less  shining,  sparsely  and  evenly  clothed  with  rather  long  recumbent 
hair-like  scales,  each  scale  longitudinally  unistriate,  grayish  in  color. 

Head  rather  strongly  convex  on  the  vertex ;  front  broadly  and  feebly 
impressed  above  the  scarcely  prominent  clypeo-f rental  junction,  a  smooth 
and  narrow  median  line  more  or  less  evident ;  punctuation  rather  coarse 
and  sparse.  Antennae  moderately  long  and  slender,  joints  less  elongate 
than  in  sequoiae  and  similar  to  those  in  squamulatus  and  its  allies,  apical 
five  joints  somewhat  stouter,  third  to  the  sixth  subequal  in  length  and 
slightly  shorter  than  the  seventh. 

Pronotum  about  a  third  wider  than  long,  widest  at  middle,  rather 
moderately  convex,  strongly  and  precipitously  declivous  at  the  sides  in 
region  of  the  apical  angles ;  apex  moderately  arcuate,  post-ocular  lobes 
distinct  and  rather  strong;  sides  moderately,  broadly  and  rather  evenly 
arcuate,  somewhat  straight  before  the  obtuse  basal  angles,  as  viewed  from 
above,  marginal  bead  distinct  in  basal  third  and  more  or  less  feeble 
anteriorly;  base  feebly  arcuate;  disk  distinctly  and  somewhat  sparsely 
punctate,  punctures  moderately  coarse,  separated  by  a  distance  equal  to 
their  diameter. 


GLYPTOSCELIS,   NEW  SPECIES  197 

Elytra  less  than  half  longer  than  wide,  rather  broadly  and  obtusely 
rounded  posteriorly ;  sides  very  feebly  arcuate,  humeri  moderately  nar- 
rowly rounded,  umbones  rather  distinct ;  base  very  slightly  emarginate  ; 
surface  strongly  transversely  convex;  punctuation  somewhat  finer  than 
on  the  pronotum,  punctures  separated  by  a  distance  equal  to  about  twice 
their  own  diameter,  still  finer  on  the  apical  declivity.  Scutellum  slightly 
transverse,  apex  feebly  angulate,  surface  with  a  few  small  punctures. 

Abdomen  rather  more  thickly  punctate  than  above. 

Legs  comparatively  short  and  rather  stout.  Anterior  tibiae  quite 
straight;  middle  and  posterior  tibiae  slightly  outwardly  curved  at  apex. 

Claws  cleft.  In  the  male  the  anterior  claws  are  cleft  to  slightly 
behind  the  middle ;  the  inner  divisions  are  one-fourth  shorter  than  the 
outer  part  and  rather  acute.  In  the  female  the  anterior  claws  are  cleft 
to  the  middle,  the  inner  divisions  are  short  and  acute,  scarcely  separated 
from  the  outer  part ;  claws  of  the  middle  and  posterior  tarsi  similar, 
more  distinctly  cleft  to  about  basal  third,  inner  portions  longer,  more 
slender  and  acute. 

Male. — Narrower.  Rather  broadly  impressed  at  middle  of  the  fifth 
ventral  segment,  this  impression  impunctate  or  more  or  less  sparsely 
punctate  and  pubescent. 

Female.  —  Broader.  Fifth  ventral  rather  moderately  concave 
throughout  the  width  and  apically  more  or  less  arcuately  deflexed. 

Measurements. — Length  (Types),  5.5-7.5  mm.;  width,  3.8-3  mm. 

Holotype,  male,  and  allotype,  female,  in  the  collection  of  the  Cali- 
fornia Academy  of  Sciences ;  paratypes  in  the  collection  of  the  Academy 
and  in  that  of  the  author. 

Type  locality. — Sacramento,  California.  Collected  by  E.  P.  Van 
Duzee,  May  28,  1918.  Beaten  from  wild  rose. 

Parvulus  somewhat  resembles  Adoxus  vitis  Fab.  in  form,  but  is  rather 
narrower  and  more  elongate.  The  individuals  vary  quite  a  little  in  form, 
some  are  shorter  and  more  robust  than  others  and  intermediates  are 
abundant.  It  is  the  smallest  species  known  to  the  author.  In  all  species 
of  Glyptoscelis  examined  there  is  a  rounded  setigerous  fovea  at  the  apical 
and  basal  angles  of  the  pronotum. 

In  pubescent,  illustris  and  sequoiae  there  is  a  circum-scutellar  depres- 
sion and  an  impression  within  the  humeral  umbones.  The  latter  impression 
is  also  present  in  parvulus. 

The  two  species  here  described  cannot  be  the  same  as  albidus  Lee., 
where  the  pubescence  is  very  easily  removable  according  to  Dr.  Horn. 
A  species  taken  in  El  Dorado  and  Calaveras  Counties  answers  the  latter 
peculiarity  very  well.  In  illustris  and  squamulatus  the  vestiture  is  quite 
persistent. 


198      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

TENEBRIONIDAE. 

Centrioptera  pectoralis,  new  species. — Form  elongate  and  sub- 
parallel.  Color  black  and  more  or  less  shining.  Pronotum  extremely 
finely  and  sparsely  punctate ;  elytra  with  distinct  and  rather  stout  muricate 
tubercles  laterally  and  on  the  apical  declivity. 

Head  less  than  moderately  convex,  fronto-epistomal  margin  quite 
evenly  arcuate,  feebly  deflexed  apical  margin  scarcely  truncate  or  slightly 
sinuate  at  middle;  punctuation  very  irregular,  leaving  large  smooth 
areas ;  punctures  moderate  in  size  and  sharp  across  the  middle  of  the 
frons  and  at  periphery  of  the  epistoma.  Antennae  slender,  joints  equal 
in  width  and  elongate,  tenth  equilaterally  triangular,  not  at  all  large, 
eleventh  small  and  short,  pointed  oval. 

Pronotum  about  a  fourth  wider  than 'long;  sides  broadly  and  evenly 
arcuate  in  anterior  two-thirds,  thence  broadly  sinuate  to  the  basal  angles 
which  are  subrectangular,  marginal  bead  moderately  coarse  and  slightly 
reflexed ;  apex  broadly  emarginate  between  the  rather  acute  and  anteriorly 
prominent  angles ;  base  broadly  and  feebly  emarginate ;  disk  moderately 
and  rather  evenly  convex,  slightly  more  declivous  at  the  apical  angles, 
scarcely  transversely  impressed  before  the  base,  punctuation  extremely 
fine  and  sparse,  scarcely  granulate  or  more  distinct  at  the  sides. 

Propleura  smooth,  opaque  with  a  few  punctures  anteriorly 

Elytra  oblong,  about  a  half  longer  than  wide,  not  strongly  convex 
centrally  but  more  strongly  and  arcuately  so  laterally;  sides  broadly  and 
feebly  arcuate,  parallel,  slightly  wider  behind  the  middle,  apex  rather 
broadly  rounded ;  disk  moderately  convex,  rather  abruptly  and  obliquely 
declivous  posteriorly ;  central  area  with  simple  punctures,  laterally  and 
on  apex  the  punctures  of  the  intervals  are  replaced  by  strong  and  moder- 
ately large  muricate  tubercles,  the  punctures  and  tubercles  serially 
arranged ;  punctures  equal  in  size,  those  of  the  striae  not  muricate  and  not 
closely  placed,  those  of  the  intervals  in  the  central  and  sutural  area  more 
distantly  spaced  and  more  or  less  granulate ;  base  scarcely  arcuate,  humeri 
obtuse  and  not  at  all  prominent. 

Parapleura  moderately  coarsely  and  sparsely  punctate. 

Sterna. — Prosternum  glabrous  laterally  and  anteriorly  along  the 
margin,  elsewhere  coarsely  rugoso-punctate ;  process  broadly  and  slightly 
impressed,  apex  sublaminately  but  briefly  mucronate,  mucro  small  and 
variable.  Mesosternum  moderately  declivous,  then  briefly  vertical,  very 
coarsely  rugoso-punctate,  narrowly  glabrous  between  the  coxae.  Meta- 
sternum  coarsely  and  sparsely  punctate. 

Abdomen  horizontal  and  not  strongly  convex,  with  large  glabrous  and 
impunctate  areas,  punctures  rather  coarse,  most  in  evidence  on  the  process, 
sides  and  fifth  segment,  a  few  scattered  across  the  other  segments. 


TENEBRIONIDAE  199 

Legs  rather  long,  moderately  strongly  and  densely  sculptured. 

Measurements. — Length,  25  mm. ;  width,  10  mm. 

Holotype,  female,  as  described  above,  in  the  collection  of  the  Cali- 
fornia Academy  of  Sciences.  An  allotype  in  my  own  collection. 

Type  locality  and  habitat. — San  Benito  Island,  off  the  coast  of  Lower 
California.  Collected  July  15,  1905,  by  F.  X.  Williams,  while  with  the 
Galapagos  Expedition. 

Pectoralis  is  distinct  by  its  parallel  form,  punctuation  of  under  sur- 
face and  other  characters ;  apparently  not  identical  with  any  described  in 
the  Biologia. 

In  spiculifera  Lee.  the  mesosternum  is  almost  declivous  anteriorly,  in 
pectoralis  it  is  strongly  so,  as  it  is  in  dulzurae  described  below.  The  meso- 
sternum is  vertical  and  emarginate  anteriorly  in  asperata  Horn,  angularis 
Horn,  variolosa  Horn,  seriata  Lee.  and  slightly  declivous  in  muricata  Lee. 
Pectoralis  somewhat  resembles  spiculifera,  with  muricata  next  in  order. 
Spiculifera  is  a  larger  and  broader  species,  with  the  abdomen  almost 
impunctate  and  the  legs  more  sparsely  and  less  coarsely  sculptured.  In 
spiculifera,  also,  the  mentum  is  transversely  cordate,  apex  feebly  emar- 
ginate, surface  slightly  convex  and  coarsely  punctate,  with  the  antennae 
less  slender  and  parallel,  with  the  ninth  and  tenth  joints  triangular,  the 
eleventh  longer.  In  pectoralis  the  mentum  is  cordate,  relatively  larger, 
somewhat  concave,  more  strongly  emarginate  at  apex,  feebly  subcarinate 
along  the  median  line,  longitudinally  and  not  strongly  impressed  laterally, 
coarsely  punctate  and  more  scabrous. 

Centrioptera  dulzurae,  new  species. — Form  elongate-oblong,  similar 
to  muricata.  Color  dull  black ;  pronotum  sculptureless,  elytra  with  muri- 
cate  tubercles  laterally  and  on  apical  declivity.  Mesosternum  declivous 
anteriorly ;  abdomen  smooth  with  a  few  scattered  punctures. 

Head  slightly  convex,  fronto-epistomal  margin  arcuate,  somewhat 
oblique  laterally ;  surface  punctate,  punctures  irregularly  and  very  sparsely 
placed,  finer  in  the  central  area.  Antennae  rather  short  with  joints  of 
equal  width,  joints  four  to  eight  suboblong,  ninth  triangular,  tenth  sub- 
quadrate,  eleventh  short  pointed  ovate.  Mentum  feebly  convex,  more 
oval  than  cordate,  slightly  transverse,  apical  emargination  not  strong; 
surface  narrowly  impunctate  at  apex,  elsewhere  rather  coarsely  and  closely 
punctured,  somewhat  longitudinally  impressed  laterally. 

Pronotum  rather  more  than  a  fourth  wider  than  long ;  sides  broadly 
and  evenly  arcuate  anteriorly,  somewhat  convergent  and  broadly  sinuate 
posteriorly  to  the  basal  angles  which  are  rectangular:  apex  truncate 
between  the  strong,  anteriorly  prominent  and  acute  angles ;  base  truncate ; 
disk  evenly  and  very  moderately  convex,  a  little  less  so  basally,  sculpture- 


200      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

less  and  smooth  with  a  few  obsolete  granules  along  the  margin,  lateral 
bead  rather  moderate  and  briefly  reflexed. 

Propleura  dull,  almost  sculptureless  and  smooth. 

Elytra  oblong-oval,  somewhat  widest  behind  the  middle,  obliquely 
declivous  posteriorly,  less  than  twice  as  long  as  wide,  almost  flattened 
dorsally;  sides  rather  broadly  rounded  and  moderately  inflexed,  feebly 
sinuate  before  the  apex  which  is  slightly  prominent ;  disk  with  ten  series 
of  punctures,  punctures  moderate  in  size,  rather  corroded,  closely  and 
slightly  irregularly  placed,  series  feebly  impressed,  intervals  perfectly  flat 
centrally  and  basally  and  with  a  single  series  of  smaller  and  widely  spaced, 
feebly  submuricate  punctures ;  laterally  and  on  the  apical  declivity  the 
intervals  are  scarcely  convex,  and  with  a  single  series  of  widely  spaced 
muricate  tubercles  which  are  scarcely  longer  than  their  width  at  base. 

Sterna. — Prosternum  smooth  and  sculptureless  anteriorly  and  later- 
ally ;  process  between  the  coxae  rather  broadly  oval,  longitudinally  im- 
pressed and  coarsely  rugoso-punctate,  apex  rounded  with  a  small  mucro 
at  tip.  Mesosternum  obsoletely  and  sparsely  punctate  laterally,  moder- 
ately declivous  and  briefly  vertical  at  middle  anteriorly,  coarsely  rugoso- 
punctate,  smooth  posteriorly  between  the  coxae.  Metasternum  smooth 
and  very  strongly  punctate. 

Parapleura  smooth,  sparsely  and  more  or  less  obsoletely  punctate. 

Abdomen  flattened  throughout  middle  third,  smooth  and  very  sparsely 
punctate;  process  oblong-quadrate,  surface  with  an  oval  and  slightly 
raised  swelling  at  middle,  otherwise  more  or  less  rugose ;  first  segment 
with  a  few  punctures  behind  the  coxae,  fourth  with  a  subapical  line  of 
punctures,  fifth  sparsely  and  evenly  punctured,  punctures  somewhat  coarse 
throughout. 

Legs  moderate  in  length ;  femora  sparsely  punctate ;  tibiae  densely 
sculptured,  each  puncture  with  a  ferruginous  seta;  tarsi  with  similarly 
colored  setae,  plantar  grooves  open. 

Male. — Narrower  and  more  parallel. 

Female. — Slightly  broader  and  a  little  inflated,  (Only  an  abdomen 
at  hand.) 

Measurements. — Length  (Male),  21  mm.;  width,  9  mm.  An  abdo- 
men of  a  female  measures  10  mm.  in  width. 

Holotype,  male,  in  my  own  collection. 

Type  locality. — Dulzura,  San  Diego  County,  California. 

Habitat. — Dulzura  and  Poway,  San  Diego  County,  California. 

Dulzurae  was  identified  for  me  as  asperata  Horn  more  than  twenty- 
five  years  ago.  For  about  twenty  years  I  had  only  an  abdomen  on  a  pin 
representing  this  species.  On  my  ranch  at  Poway,  where  I  lived  for 
eleven  years,  I  took  only  one  living  specimen,  but  the  dead  bodies  of  this 


SCHIZILLUS    LATICEPS    HORN  201 

beetle  were  very  abundant  in  the  nests  of  wood  rats.  Dulzurae  is  noc- 
turnal in  habits.  Having  secured  but  one  living  specimen  in  eleven  years 
does  not  speak  well  for  my  ability  as  a  collector ;  unfortunately  I  was  more 
interested  in  farming  at  that  time  than  in  worrying  over  beetles. 

In  appearance  dulzurae  resembles  pectoralis,  which  is  a  larger  species  ; 
next  in  order  it  resembles  muricata  Lee.,  which  is  a  more  or  less  shining 
species,  a  moderate  series  of  which  is  before  me.  In  muricata  the 
propleura  are  distinctly  punctate,  the  abdomen  is  quite  coarsely  and  much 
more  abundantly  punctate  and  the  parapleura  are  more  densely  and 
strongly  punctured.  The  pronotum  of  muricata  is  impressed  and 
punctate-granulate  along  the  side  margins.  Pectoralis  is  more  elongate 
and  more  strongly  sculptured  beneath.  The  apparent  sequence  of  species 
should  be  spiculifera,  pectoralis,  muricata  and  dulzurae;  then  follow 
angularis,  asperata,  variolosa  and  seriata.  Infausta  and  utahensis  I  have 
not  seen.  This  arrangement  agrees  with  that  given  in  Henshaw's  List. 

Schizillus  Horn. 

In  1874,  Dr.  Geo.  H.  Horn  created  the  above  genus  for  laticeps,  a 
member  of  the  Tribe  Cryptoglossini,  collected  on  the  Mohave  Desert, 
California,  by  Mr.  Crotch.  In  1913,  Mr.  J.  R.  Slevin  took  a  considerable 
series  at  Hesperia,  California,  which  are  in  the  collection  of  the  California 
Academy  of  Sciences. 

In  1908,  Mr.  F.  W.  Nunenmacher  took  three  interesting  specimens 
at  Goldfield,  Nevada.  Two  of  these  look  like  a  modified  laticeps,  the  other 
somewhat  resembles  the  smooth  Cryptoglossa  laevis.  For  more  than 
twenty-eight  years  I  have  had  a  specimen  that  was  unique  as  far  as  my 
collection  was  concerned.  During  this  time  it  was  labeled  laticeps,  but  it 
undoubtedly  represents  a  new  species.  From  the  material  before  me  it  is 
evident  that  three  new  species  are  to  be  recognized. 

The  salient  generic  characters  of  Schizillus  are  the  completely  divided 
eyes,  the  transverse  mentum,  the  broader  genae  and  the  front  is  hemi- 
hexagonal.  The  antennae  are  eleven-jointed,  first  joint  stout  and 
invisible  from  above,  the  terminal  joint  oval  and  smaller  than  the  pre- 
ceding. In  regard  to  the  size  of  the  eleventh  joint,  I  must  differ  from 
Dr.  Horn,  for  it  is  smaller  than  the  preceding.  The  broad  head  is  a 
striking  character  when  compared  with  our  Centriopterae.  Dr.  Horn's 
description  of  the  type  species  may  be  modified  as  follows : 

Schizillus  laticeps  Horn. — Form  elongate  oblong-oval  and  moderately 
robust.  Color  black  and  subopaque. 

Head  broad,  front  hemi-hexagonal  and  moderately  convex,  sparsely 
punctured,  very  broadly  impressed  between  the  eyes  along  the  frontal 


202      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

suture,  punctures  moderately  small  and  densest  along  the  apical  margin, 
neck  densely  and  finely  granulate.  Antennae  rather  stout  and  parallel,  the 
terminal  four  joints  slightly  narrower  and  less  stout  than  the  preceding 
joints. 

Pronotum  broader  than  long;  apex  moderately  deeply  emarginate, 
slightly  broader  than  the  base ;  sides  moderately  arcuate,  sinuate  near  the 
base,  hind  angles  rather  less  than  rectangular ;  base  broadly  emarginate ; 
surface  moderately  convex,  sparsely  punctured  and  subopaque  and  with  a 
more  or  less  deep,  transverse,  ante-basal  impression;  apical  angles  quite 
broad,  anteriorly  prominent  and  subobtuse. 

Propleura  sparsely  punctate,  punctures  subobsolete,  many  rugulae 
more  or  less  evident. 

Elytra  oblong-oval,  not  wider  at  base  than  the  pronotum,  widest  at 
middle,  sides  moderately  arcuate;  surface  moderately  convex,  with  rows 
of  punctures  on  the  disk  which  become  irregular  and  submuricate  at  the 
sides  and  apex,  intervals  flat  and  submuricately  punctate ;  serial  punctures 
a  little  larger  than  the  interstitial ;  disk  almost  vertically  declivous 
posteriorly. 

Parapleura  irregularly  and  sparsely  punctate. 

Sterna. — Prosternum  sparsely  punctate,  more  or  less  impunctate 
within  the  oblique  sutures ;  process  slightly  produced,  subevenly  oval, 
margin  prominent,  surface  impressed  and  sparsely  punctate  and  with  a 
few  rugae.  Mesosternum  moderately  declivous,  quite  deeply  impressed, 
surface  smooth  and  narrowly  punctate  along  the  periphery.  Metasternum 
very  short  and  punctato-rugose. 

Abdomen  sparsely  and  very  obsoletely  punctate;  intercoxal  process 
rugose,  fourth  and  fifth  segments  distinctly  punctate  apically,  the  former 
glabrous  in  about  basal  half,  the  latter  in  basal  third;  first  suture  per- 
fectly straight,  the  third  and  fourth  quite  strongly  arcuate. 

Legs  rather  short.  'Femora  rather  sparsely  punctate,  tibial  grooves 
distinctly  and  rather  more  densely  punctured;  tibiae  quite  densely 
muricato-punctate ;  tarsi  stout  and  similar  in  the  sexes,  plantar  grooves 
open,  the  setae  ferruginous. 

Male. — Narrower.  Abdomen  feebly  oblique  to  the  'sterna  and 
slightly  more  flattened  between  the  metacoxae. 

Female. — Broader.  Abdomen  perfectly  horizontal  and  slightly  more 
convex.  Elytra  more  broadly  oblong-oval. 

Measurements. — Length,  21-25  mm. ;  width,  9-11  mm. 

Type  in  the  Horn  collection  at  the  Philadelphia  Academy  of  Sciences. 

Type  locality. — Mohave  Desert,  California. 

Habitat. — Hesperia  and  the  Mohave  Desert,  California. 


SCHIZILLUS    LATICEPS    HORN  203 

In  laticeps  the  elytral  sculpturing  resembles  that  observed  in  Eleodes 
consobrina  Lee. ;  the  epipleura  are  sparsely  and  very  obsoletely  punctate 
and  smooth.  It  is  a  large  and  more  elongate  species  with  abdomen  very 
smoothly  sculptured. 

Schizillus  convexus,  new  species. — Form  oblong-oval.  Color  black  and 
more  or  less  subopaque. 

Head  broadly  and  strongly  transversely  impressed  between  the 
antennae,  vertex  convex  and  declivous;  fronto-epistomal  margin  some- 
what hemi-hexagonal,  almost  evenly  arcuate ;  surface  more  or  less  densely 
and  irregularly  punctate  on  the  epistoma,  front  sparsely  punctured,  punc- 
tures moderately  coarse.  Antennae  slender,  joints  nearly  of  equal  width 
throughout,  last  three  joints  almost  narrower  than  the  preceding  joints, 
ninth  and  tenth  longer  than  wide  and  subtriangular,  eleventh  smaller  and 
pointed-oval,  a  little  longer  than  wide ;  apical  joints  with  a  number  of 
rather  long  tactile  hairs. 

Pronotum  about  a  third  wider  than  long;  apex  slightly  broader  than 
the  base,  moderately  deeply  emarginate,  margin  rather  straight  and 
oblique  within  the  angles ;  sides  moderately  arcuate  in  anterior  three- 
fourths,  sinuate  before  the  base,  marginal  bead  coarse  and  reflexed  only 
near  the  angles ;  base  broadly  and  not  strongly  emarginate,  beaded ;  basal 
angles  subrectangular  ;  disk  smooth  and  moderately  convex,  more  strongly 
so  anteriorly  and  centrally,  moderately  declivous  at  the  apical  angles, 
somewhat  impressed  within  the  basal  angles,  more  or  less  obsoletely  to 
distinctly  punctate,  punctures  rather  coarse  and  more  distinct  laterally; 
transverse  ante-basal  impression  short,  not  strongly  defined  and  slightly 
more  impressed  at  middle  and  more  strongly  so  within  the  basal  angles. 

Propleura  rather  coarsely,  sparsely  but  distinctly  punctate  and 
rugose. 

Elytra  oblong-oval,  widest  a  little  behind  the  middle  (in  type),  base 
scarcely  wider  than  the  pronotal  base ;  surface  moderately  convex,  strongly 
rounded  at  the  sides,  rows  of  punctures  not  distinct,  muricately  punctate 
throughout,  more  strongly  so  on  the  sides  and  apical  declivity,  the  latter 
rather  abruptly  oblique ;  muricate  punctures  shining  at  tip  and  subequal 
throughout ;  apex  somewhat  lobed  and  rather  broadly  rounded. 

Parapleura. — Mesosternal  side-pieces  distinctly  and  densely  punctate ; 
metasternal  side  pieces  sparsely  punctured,  punctures  larger. 

Sterna. — Prosternum  rather  coarsely  and  quite  densely  punctate 
throughout;  process  slightly  produced,  rather  broadly  rounded  at  tip, 
marginal  bead  coarse  and  raised ;  surface  depressed,  coarsely  and  densely 
punctate.  Mesosternum  closely  punctate,  declivous,  deeply  impressed  at 
middle,  impression  glabrous.  Metasternum  very  short,  densely  punctate 
and  rugose. 


204      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Abdomen  more  or  less  closely  and  moderately  coarsely  punctate, 
intercoxal  process  rugose  and  smooth  at  middle;  segments  more  or  less 
transversely  rugose  along  the  sides,  second  segment  more  or  less  glabrous 
in  middle  third,  third  quite  glabrous  at  middle,  fourth  and  fifth  rather 
more  densely  punctate,  fourth  more  or  less  glabrous  along  the  base ;  first 
suture  straight,  third  and  fourth  strongly  arcuate. 

Legs  rather  slender,  posterior  tibiae  slightly  inwardly  arcuate ;  femora 
densely  punctate  throughout  including  the  tibial  grooves,  punctures  smaller 
than  those  of  the  abdomen ;  tibiae  densely  muricato-punctate ;  tarsi  similar 
in  the  sexes,  rather  stout,  plantar  grooves  open ;  terminal  joints  of  all  of 
the  tarsi  fringed  with  rather  long  and  closely-placed  ferruginous  setae. 

Male  unknown. 

Female. — Abdomen  convex  and  horizontal.  Elytra  oblong  subovate 
and  moderately  broad. 

Measurement. — Length  (Type),  23  mm. ;  width,  10  mm. 

Holotype,  female,  in  my  own  collection.  A  paratype  in  the  collection 
of  Mr.  F.  W.  Nunenmacher. 

Type  locality. — Goldfield,  Esmeralda  County,  Nevada.  Collected 
June  16,  1908.  Two  females  studied. 

Convexus,  as  compared  with  'laticeps,  is  more  convex  and  not  so 
broad,  the  body  beneath  is  much  more  densely  and  strongly  punctate,  the 
pronotum  is  more  distinctly  punctate,  especially  within  the  angles,  the 
mentum  is  less  transverse,  densely  punctured,  with  the  sides  broadly 
arcuate  and  the  surface  at  apex  impressed  so  as  to  make  the  margin 
appear  slightly  emarginate. 

Schizillus  nunenmacheri,  new  species. — Form  elongate,  oblong-sub- 
ovate.  Color  dull  black,  more  or  less  opaque.  Subobsoletely  sculptured. 

Head  very  broadly  and  transversely  but  not  strongly  impressed 
between  the  antennae  and  along  the  sides  before  the  eyes ;  central  area  of 
the  epistoma  slightly  prominent  and  convex ;  fronto-epistomal  margin 
rather  hemi-hexagonal ;  surface  smooth,  obsoletely  punctate,  except  on  the 
epistoma  along  the  margin  where  the  punctures  are  more  or  less  distinct. 
Antennae  quite  slender,  third  joint  very  elongate,  about  equal  to  the  next 
four  taken  together,  joints  four  to  eleven  rather  narrower  than  the  third, 
about  twice  as  long  as  wide,  tenth  slightly  narrower,  eleventh  smaller, 
pointed-oval  and  about  as  long  as  wide. 

Pronotum  almost  a  half  wider  than  long;  apex  rather  deeply  and 
broadly  emarginate,  apical  angles  subacute  and  anteriorly  prominent; 
sides  feebly  arcuate  in  anterior  one-half,  thence  almost  straight  and 
moderately  convergent  to  the  basal  angles,  marginal  bead  very  fine  to 
subobsolete;  base  broadly  and  feebly  emarginate;  basal  angles  sub- 


SCHIZILLUS    LATICEPS    HORN  205 

rectangular;  disk  sculptureless,  smooth,  almost  without  impressions,  cen- 
tral area  anteriorly  rather  strongly  convex,  elsewhere  moderately  so, 
gradually  declivous  at  the  apical  angles. 

Parapleura  smooth  and  opaque,  with  few  rugae  on  the  coxal  con- 
vexities. 

Elytra  about  a  half  longer  than  wide,  suboblong-oval,  widest  at  about 
the  middle ;  base  feebly  arcuate,  quite  equal  to  the  pronotal  base ;  disk 
moderately  convex,  more  strongly  and  rather  broadly  rounded  at  the  sides, 
obliquely  declivous  posteriorly;  apical  lobe  moderate  and  not  broadly 
rounded ;  surface  smooth  and  dull,  striae  obsolete  or  very  faintly  evident 
when  viewed  obliquely,  interstices  with  a  row  of  rather  small  muricate 
tubercles ;  these  are  obsolete  along  the  suture,  becoming  gradually  stronger 
laterally  and  on  the  apical  declivity. 

Epipleura  smooth  and  without  sculpturing. 

Sterna. — Prosternum  obsoletely  rugose  anteriorly  and  on  the  process, 
the  latter  a  little  produced,  suboval  with  the  sides  converging  slightly 
behind  the  coxae,  tip  rather  broadly  rounded,  marginal  bead  not  evident ; 
surface  longitudinally  impressed  along  the  middle,  with  few  punctures 
along  the  margin.  Mesosternum  not  sculptured,  declivous  anteriorly, 
broadly  impressed  at  middle,  sides  prominent  and  subtuberculate.  Meta- 
sternum  rugose  at  middle,  smooth  laterally. 

Parasterna  sculptureless,  or  with  a  few  fine  obsolete  punctures. 

Abdomen  obsoletely  sculptured;  a  few  very  fine  punctules  more  or 
less  evident,  few  coarse  punctures  behind  the  coxae;  fifth  segment  with 
very  small  indistinct  punctures. 

Legs  rather  slender,  anterior  pair  apparently  a  little  stouter.  Femora 
and  tibiae  rather  densely  but  not  strongly  sculptured  throughout  including 
the  tibial  grooves,  the  former  punctate,  the  latter  muricato-punctate. 
Tarsi  without  doubt  similar  in  the  sexes,  rather  stout,  plantar  grooves 
open,  setae  ferruginous;  terminal  joints  of  all  the  tarsi  fringed  with 
closely  placed  ferruginous  setae. 

Male  unknown. 

Female. — Abdomen  convex  and  horizontal. 

Measurements — Length,  18.2  mm. ;  width,  8  mm. 

Holotype  a  female  in  my  collection.  Collected  on  Aug.  14,  1908,  by 
Mr.  F.  W.  Nunenmacher. 

Type  locality  and  habitat. — Goldfield,  Esmeralda  County,  Nevada. 

Nunenmacheri  is  distinct  in  its  obsolete  sculpturing,  long  third  an- 
tennal  joint  and  less  rounded  sides  of  the  pronotum.  The  mentum  is 
smooth  and  partly  hidden  by  dried  regurgitated  ingesta.  This  species  is 
suggestive  of  Cryptoglossa  laevis  on  account  of  its  smooth  integuments, 
with  the  exception  of  the  elytra ;  it  is  more  elongate  and  less  robust,  the 


206      NEW  SPECIES  MELYR1DAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

mesosternum  is  different  and  the  eleventh  joint  of  the  antennae  is  oval  as 
in  Centrioptera  while  in  laevis  it  is  short  and  truncate.  The  form  is 
similar  in  nunenmacheri  and  convexus,  but  the  latter  is  larger. 

Schizillus  lomae,  new  species. — Form  oblong-oval,  moderately  broad 
and  rather  robust,  elytra  slightly  flattened  on  the  disk.  Color  black, 
feebly  shining,  somewhat  alutaceous  anteriorly. 

Head  moderately  broad,  feebly  and  broadly  impressed  between  the 
antennae,  epistoma  very  feebly  convex  at  middle ;  surface  smooth, 
sparsely  punctate ;  punctures  small  and  not  strong,  densely  placed  along 
the  fronto-epistomal  margin  which  is  hemi-hexagonal.  Antennae  slender, 
joints  of  equal  width,  third  joint  a  little  longer  than  the  fourth,  the  latter 
a  little  longer  than  the  fifth ;  fifth,  sixth  and  seventh  nearly  subequal,  ninth 
and  tenth  somewhat  compressed,  tenth  triangulo-oval,  eleventh  oval,  about 
as  long  as  wide,  pointed  at  middle  of  tip. 

Pronotum  about  a  third  wider  than  long,  transverse ;  apex  deeply 
emarginate,  bottom  of  the  emargination  almost  transverse  in  the  middle 
two-fourths,  thence  oblique  to  the  apical  angles,  which  are  subacute, 
broad  and  anteriorly  prominent,  marginal  bead  present  within  the  angles ; 
sides  strongly,  broadly  and  evenly  arcuate  in  the  anterior  four-fifths,  thence 
sinuate  and  parallel  to  the  basal  angles,  marginal  bead  rather  coarse  and 
somewhat  reflexed ;  base  broadly  and  arcuately  emarginate,  bead  coarse ; 
basal  angles  less  than  a  right  angle,  blunt  and  somewhat  prominent  pos- 
teriorly, marginal  bead  strong  about  the  angles ;  surface  smooth  and 
obsoletely  sculptured,  moderately  strongly  convex  in  the  central  area, 
especially  anteriorly,  less  so  posteriorly,  rather  broadly  impressed  laterally, 
with  a  gutter  posteriorly  which  is  continuous  with  the  short  ante-basal 
transverse  impression,  the  latter  moderately  strong;  slightly  impressed 
within  the  apical  angles  and  strongly  impressed  within  the  basal  angles. 

Propleura  smooth,  with  a  submarginal  line  of  short  rugulae. 

Elytra  broadly  oval,  about  a  fourth  longer  than  wide,  dorsum  rather 
feebly  convex,  strongly  but  not  broadly  rounded  at  the  sides,  apical 
declivity  almost  vertical,  apical  lobe  short  and  broadly  rounded ;  humeri 
obsolete,  base  equal  to  the  pronotal  base ;  scutellum  very  small  and  tri- 
angular; surface  sculptured  with  rather  close  series  of  punctures  which 
are  simple  in  the  central  area  of  the  disk,  becoming  muricate  and  then 
muricato-rugose  on  the  lateral  and  apical  declivities  ;  strial  punctures  more 
closely  placed  and  slightly  larger,  the  interstitial  series  a  little  smaller  and 
more  widely  spaced. 

Epipleura  rather  wide  and  obsoletely  sculptured. 

Parapleura. — Mesosternal  pieces  rather  densely  punctured ;  meta- 
sternal  pieces  sparsely  punctate,  punctures  a  little  larger,  surface  smooth. 


SCHIZILLUS    LATICEPS    HORN  207 

Sterna. — Prosternum  not  strongly  punctate  in  front  of  the  coxae, 
rather  transversely  glabrous  at  middle.  Process  slightly  produced,  oval 
between  the  coxae  with  the  sides  arcuately  converging  posteriorly  to  the 
obtuse  tip ;  surface  broadly  and  feebly  impressed,  rather  densely  punctate, 
bead  not  distinct.  Mesosternum  arcuately  declivous  anteriorly,  rather 
deeply  impressed  in  middle  third,  sides  coarsely  punctate.  Metasternum 
coarsely  and  irregularly  rugose,  sides  sparsely  punctate  and  rather  smooth. 

Abdomen  not  strongly  convex,  slightly  oblique  to  the  sterna;  surface 
smooth,  with  widely  scattered  fine  punctures  on  the  first  three  segments ; 
the  punctures  are  slightly  stronger  on  the  apical  half  of  the  third  segment ; 
intercoxal  process  rugose ;  fourth  and  fifth  segments  more  or  less  densely 
punctate,  punctures  small ;  first  suture  almost  straight,  third  and  fourth 
strongly  arcuate,  especially  at  the  middle. 

Legs  rather  long  and  moderately  slender,  anterior  pair  a  little  stouter, 
rather  densely  sculptured,  femora  punctate,  tibiae'  not  coarsely  muricato- 
punctate ;  tarsi  stout,  plantar  grooves  open,  setae  ferruginous ;  terminal 
joints  of  all  the  tarsi  fringed  with  rather  long  and  closely-placed  ferru- 
ginous setae. 

Male. — Abdomen  not  strongly  convex  and  slightly  oblique  to  the 
sterna. 

Female. — Unknown. 

Measurements. — Length,  20  mm.;  width,  10  mm.;  elytral  length, 
13  mm. 

Holotype,  male,  in  my  collection.     Collected  by  Mr.  O.  N.  Sanford. 

Type  locality  and  habitat. — Point  Loma,  San  Diego,  California. 

Lomae  is  very  distinct  from  the  other  species.  The  mentum  is 
cordato-oval,  slightly  transverse,  apex  slightly  emarginate,  lobes  arcuately 
rounded  into  the  broadly  arcuate  sides ;  surface  distinctly  but  not  strongly 
convex,  closely  punctured,  punctures  moderate  in  size. 

The  scutellum  in  the  Tenebrionidae  as  a  rule  is  a  very  unreliable  and 
variable  structure  in  any  well  defined  species.  In  Schisillus  it  enters 
between  the  elytra  but  a  short  distance.  The  scutellum  is  more  or  less 
triangular,  short  and  more  transverse  in  laticeps,  convexus  and  nunen- 
macheri.  In  the  type  of  lomae  it  is  small  and  equilaterally  triangular. 

The  following  table  will  serve  to  separate  the  species  of  the  genus 
Schizillus : 

Humeri  obsolete,  elytra  broadly  oval lomae,  n.  sp. 

Humeri  more  or  less  distinct,  elytra  oblong  oval 1 

1.  Third  antennal  joint  very  elongate,  about  equal  in  length  to  the  four  fol- 

lowing joints imnenmacheri,  n.  sp. 

-.    Third  joint  shorter  than  the  next  two  taken  together 2 

2.  Abdomen  very  finely  and  sparsely  punctate - laticeps  Lee. 


208      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

-.    Abdomen  very  distinctly,  rather  closely  punctured,  punctures  moderate  in 

size convexus,  n.  sp. 

The  following  antennal  differences  may  be  noted : 

Third  joint  a  little  longer  than  the  fourth,  the  latter  relatively  longer  than  in 

laticeps  _....,. lomae 

Third  joint  about  equal  to  the  next  four  taken  together,  these  being  subequal 

in  length  and  about  a  third  longer  than  wide nunenmacheri 

Third  joint  about  twice  as  long  as  wide,  distinctly  shorter  than  the  next  two 

taken  together;  fourth,  fifth  and  sixth  subequal  in  length laticeps 

Third  joint  distinctly  twice  as  long  as  wide,  fourth  longer  than  the  fifth convexus 

Euschides  lecontei  Horn. — This  species  inhabits  the  valleys  and  foot- 
hills of  the  southern  San  Joaquin  Valley.  To  what  altitude  in  the  foot- 
hills is  not  definitely  known.  Series  are  before  me  that  were  collected 
near  Los  Angeles  and  at  Bakersfield.  There  is  a  noticeable  difference 
between  the  extremes  of  the  two  series.  In  the  Bakersfield  specimens  the 
males  are  comparatively  slender  and  quite  parallel,  the  elytra  are  dis- 
tinctly narrower  than  the  pronotum ;  in  the  female  the  elytra  are  broader 
and  distinctly  wider  than  the  pronotum,  with  the  sides  moderately  arcuate ; 
in  both  sexes  the  dorsum  is  less  convex  than  in  the  Los  Angeles  speci- 
mens. In  the  latter  the  males  are  less  narrow  and  parallel,  the  elytra 
being  at  least  as  wide  as  the  pronotum,  with  the  sides  broadly  and  more 
strongly  arcuate  than  in  the  Bakersfield  males ;  in  the  females  there  is 
often  present  a  short  costa  between  the  margin  and  the  second  discal, 
these  costae  are  not  strong  and  are  usually  obsolescent;  both  sexes  are 
more  robust  and  the  dorsum  more  convex. 

In  both  series  the  sculpturing  is  similar  and  the  variations  are  analo- 
gous in  every  way.  The  surface  lustre  is  somewhat  duller  in  the  Bakers- 
field  specimens.  The  characters  appertaining  to  the  two  series  as  men- 
tioned above,  gradate  completely  one  into  the  other.  The  largest  speci- 
men mentioned  by  Casey  measured  20  mm. 

In  the  collection  of  the  California  Academy  of  Sciences  is  a  specimen, 
as  well  as  one  in  my  own,  that  was  collected  in  the  Kings  River  Canyon, 
California,  both  taken  by  Mr.  Frank  Daggett ;  these  two  specimens  are 
giants  as  compared  to  the  other  specimens  seen  by  me.  They  measure 
as  follows :  Length,  24-25  mm. ;  width,  10-10.3  mm.  In  these  the  discal 
costae  become  obsolescent  before  becoming  coalescent  on  the  apical 
declivity  and  there  is  no  evidence  of  a  third  costa.  The  pronotum  is 
transverse,  disk  more  convex  and  less  deplanate  at  the  sides,  the  central 
area  is  smooth,  sparsely  punctate,  punctures  smaller,  less  strong,  becoming 
coarser  laterally  and  on  the  deplanate  area  more  or  less  punctato-rugose, 
and  the  margin  is  less  reflexed  than  in  the  two  series  mentioned  above ;  the 
base  is  nearly  arcuate,  scarcely  bisinuate;  the  head  has  a  large  cordate 
depression.  The  legs  are  relatively  stouter  than  in  the  Los  Angeles  and 


CONIONTIS  SANTAROSAE  209 

Bakersfield  specimens ;  in  the  former  they  are  stouter  than  in  the  latter, 
where  they  are  quite  slender.  In  both  the  Bakersfield  and  Los  Angeles 
specimens  the  antennae  are  slender  and  the  last  three  joints  form  a  club ; 
in  those  from  Kings  River  Canyon  the  antennae  are  stouter  and  gradually 
incrassate  and  the  under  surface  of  the  body  is  more  coarsely  and  densely 
punctate.  For  this  large  phase  I  propose  the  name  gigantea,  new  sub- 
species. Among  the  specimens  referred  to  these  three  series  there  are  the 
usual  individual  variations  observed  in  other  species,  and,  when  these  are 
arranged  in  a  block  system,  the  gradations  are  complete.  These  forms 
are  the  result  of  geographical  position  and  environment  which  produce 
such  modifications  and  reveal  the  plasticity  and  adaptability  of  the 
organisms. 

Coniontis  santarosae,  new  species. — Form  oblong-oval,  about  twice 
as  long  as  wide,  rather  more  than  moderately  convex.  Color  black,  tibiae 
and  tarsi  piceous,  surface  smooth  and  shining;  pubescence  microscopical 
above,  longer  and  more  abundant  beneath. 

Head  very  finely  and  sparsely  punctate,  punctures  stronger  and  more 
numerous  on  the  epistoma ;  frontal  suture  distinct ;  epistomal  sinuation 
moderate  and  arcuately  reentrant.  Antennae  shorter  than  the  length  of 
the  pronotum,  three  terminal  joints  subequal  in  size  and  moderately  com- 
pressed, scarcely  wider  than  the  preceding  joints. 

Pronotum  almost  twice  as  wide  as  long,  evenly  arcuate  from  side  to 
side,  more  strongly  arcuate  and  declivous  at  the  apical  angles ;  sides  not 
strongly  convergent  anteriorly,  feebly  arcuate,  more  strongly  so  in  apical 
one-half;  marginal  bead  fine;  apex  arcuate  in  circular  arc,  almost  trun- 
cate when  viewed  from  above,  finely  beaded,  but  less  so  at  middle ;  base 
almost  truncate,  very  feebly  sinuate  laterally,  angles  scarcely  more  promi- 
nent posteriorly  than  the  broad  and  very  feebly  arcuate  median  lobe ;  basal 
angles  subrectangular ;  apical  angles  obtusely  and  narrowly  rounded ;  disk 
with  sparse  and  extremely  fine  punctules,  each  with  a  microscopical  hair. 

Elytra  oblong,  about  a  third  longer  than  wide,  evenly  arcuate  trans- 
versely, more  gradually  so  on  the  apical  declivity;  surface  smooth  and 
polished,  sometimes  very  finely  wrinkled,  feeble  grooves  and  costae  some- 
times noticeable ;  each  puncture  with  a  microscopical  hair. 

Prosternum  feebly  and  sparsely  punctured,  with  numerous  pseudo- 
punctules  scattered  throughout ;  process  finely  margined  on  apex  or  not. 

Abdomen  very  sparsely  and  finely  punctulate. 

Male. — Somewhat  narrower.  Pronotum  relatively  broader.  Ab- 
domen less  convex. 

Female.— Broader.  Pronotum  relatively  narrower.  Abdomen  more 
convex. 


210      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Measurements. — Length  (Types),  12.0  mm.;  width,  6.0-6.5  mm. 

A  series  of  fifty-two  specimens  studied. 

Variations. — These  include  those  of  size,  slight  cuneateness  in  some 
males,  feebler  and  stronger  punctuation  and  rugulosity  of  the  elytral  sur- 
face, margining  of  the  prosternal  process  not  constant ;  posterior  canthi 
of  the  eyes  not  always  the  most  prominent,  occasionally  stronger  on  one 
side  and  feebler  on  the  other. 

Extremes  in  size  measure  as  follows:  Length  (Male),  9.0  mm.; 
(Female),  12.5  mm.;  width,  4.0-6.5  mm. 

Holotype,  male,  and  allotype,  female,  in  the  collection  of  the  Cali- 
fornia Academy  of  Sciences ;  paratypes  in  the  same  and  the  author's  col- 
lection. 

Type  locality. — Santa  Rosa  Island,  off  the  coast  of  Southern  Cali- 
fornia. Collected  by  Mr.  Van  Duzee,  May  20,  1919. 

Habitat. — Santa  Rosa  and  San  Miguel  Islands.  Mr.  Van  Duzee 
says:  "Found  under  dry  'cow-chips'  near  the  entrance  to  the  pier  on 
Santa  Rosa  Island,  where  they  seemed  to  be  abundant." 

Santarosae  was  at  first  referred  to  Coniontides  infinitimus  Casey,  as 
that  species  according  to  its  author  is  more  elongate  than  latus  or  insularis. 
The  habitat  of  infiinitimus  is  uncertain  but  Casey  believes  it  to  be  the 
Island  of  Santa  Rosa.  The  form  of  the  pronotal  base  in  santarosae  for- 
bids any  association  with  the  Coniontides.  Its  general  facies  associates 
it  with  the  viatica  group  while  the  measurements  suggest  the  robusta 
group.  These  groups  are  arbitrary  and  can  only  be  used  as  a  convenience. 

The  mentum  in  santarosae  is  transverse,  lobes  obtusely  angulate  at 
apex,  sides  arcuate ;  apex  broadly  sinuate  in  the  middle  three-fifths,  sinua- 
tion  more  or  less  moderate  in  depth  and  arcuately  reentrant ;  surface  feebly 
convex,  slightly  asperate  and  irregular  on  the  lobes,  more  or  less  im- 
pressed at  base  within  the  broad  margin  and  sparsely  and  more  or  less 
regularly  punctate,  the  punctures  shallow  and  each  with  a  short  yellow 
hair. 

Coniontis  musculus  Blais. — One  of  the  most  important  things  relat- 
ing to  the  study  and  knowledge  of  organisms  is  habitat.  Habitat,  when 
known,  makes  it  possible  to  correlate  the  effect  of  geographical  position 
and  environment  on  the  organisms  of  different  regions.  In  the  majority 
of  cases  when  that  knowledge  is  lacking  it  is  folly  to  describe  a  species  or 
race,  all  data  accredited  to  such  a  species  is  presumptive  or  a  pure  pos- 
tulation  and  without  scientific  value.  Especially  is  this  true  in  cases  of 
organisms  that  are  particularly  variable  as  regards  individual  form  and 
sculpturing,  as  in  such  genera  as  Coniontis,  Eleodes,  Pterostichus  and 
many  others. 


CONIONTIDES   CASEY  211 

In  the  collection  of  the  California  Academy  of  Sciences  there  are 
specimens  of  a  species  of  Coniontis  which  must  be  referred  to  musculus 
Blais.  These  specimens  were  collected  in  the  Kings  River  Canyon  by 
Mr.  Frank  Daggett,  no  altitude  is  given  and  this  is  a  stumbling  block  to  a 
correct  understanding  of  these  specimens.  Musculus  inhabits  the  San 
Joaquin  Valley  as  far  south  as  Tehachapi,  and  westward  and  eastward 
into  the  foothills,  to  just  what  altitude  it  attains  is  not  known.  Musculus 
appears  to  be  exceedingly  abundant  in  Ventura  County,  especially  in  the 
vicinity  of  Oxnard.  The  specimens  from  this  latter  locality  show  some 
deviation  from  the  typical  form  from  the  more  northern  limit  of  its  dis- 
tribution. So  it  is  with  the  specimens  from  the  Kings  River  Canyon; 
these  are  relatively  narrower,  and  therefore  less  robust  than  the  true 
musculus.  If  all  of  the  specimens  that  I  have  seen  referable  to  musculus 
could  be  arranged  in  a  block  system,  the  observable  differences  would 
become  evanescent,  for  a  few  individuals  from  each  locality  would  resemble 
similar  ones  from  other  places.  It  is  often  the  general  habitus  or  facies 
in  the  broad  sense  that  should  guide  in  the  limitation  of  species  in  such 
genera  as  Coniontis,  Eleodes  and  others. 

Genus  Coniontides  Casey. — A  series  of  sixty-nine  specimens  in  this 
genus  are  before  me.  They  were  collected  on  the  different  islands  off 
the  coast  of  Southern  California,  namely:  San  Clemente,  Santa  Rosa, 
Santa  Barbara,  Santa  Cruz,  Anacapa,  Prince  and  San  Nicolas;  on  the 
mainland  near  Los  Angeles  a  small  series  was  taken  by  Dr.  E.  C.  Van 
Dyke. 

The  character  that  gives  Coniontides  any  claim  to  generic  standing,  is 
"the  marked  prolongation  backward  of  the  thoracic  angles,  recalling  the 
Eusatti,  and  in  every  way  similar  to  the  form  there  prevailing"  (Casey), 
and  in  the  insular  habitat ;  Dr.  Van  Dyke's  series,  however,  shows  that  the 
species  are  not  entirely  confined  to  the  islands.  Coniontides  has  as  much 
claim  to  generic  standing  as  has  Coniontellus ;  in  the  latter  the  eyes  are 
completely  divided  by  the  sides  of  the  head,  otherwise  it  is  similar  to 
Coniontis.  In  Coelus  and  Coelotaxis  the  first  joint  of  the  anterior  tarsi  is 
prolonged  beneath  the  second,  but  in  the  former  it  is  much  longer.  I  see 
no  reason  why  Coniontides  should  not  remain  as  a  valid  genus. 

Coniontides  latus  Lee.— As  usual  in  a  large  series,  there  is  great 
variation  in  size  and  sculpturing.  The  condition  of  cuneateness  possessed 
by  many  males  is  not  a  group,  a  generic,  nor  a  specific  character.  It  is 
observed  in  all  species  and  races  of  Coniontis.  Body  form  or  outline 
varies  greatly  and  between  the  extremes  all  degrees  of  intermediates  are 
found. 

The  connectants  invalidate  any  particular  value  assigned  to  a  given 
form.  The  prosternal  process  is  usually  unmargined,  some  specimens, 


212      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

however,  have  it  margined  and  all  degrees  exist  between  the  extremes. 
The  mentum  presents  great  variation  as  regards  sculpturing;  some  speci- 
mens have  the  lobes  quite  deeply  concave  or  impressed  with  the  surface 
subcarinate  in  the  median  line  and  rather  coarsely  punctured  throughout ; 
others  have  the  surface  feebly  convex,  not  at  all  impressed,  but  punctate 
as  indicated  above,  or  more  or  less  glabrous  at  the  apical  sinuation.  All 
intermediate  degrees  are  at  hand.  Punctuation  is  the  only  character 
available  for  subdividing  the  species  into  races.  Here  the  separation  is 
artificial,  arbitrary  and  opinionative. 

In  latus  the  elytral  punctures  are  almost  as  coarse  as  are  those  of  the 
pronotum  and  the  surface  lustre  is  dull.  Specimens  collected  Novem- 
ber 3,  1916,  on  the  Island  of  San  Clemente  are  before  me. 

Clementinus  Casey  is  only  an  individual  variation,  a  certain  number 
of  specimens  can  be  selected  to  constitute  a  group.  This  phase  is  identical 
with  latus  and  the  name  falls  in  synonymy. 

In  the  race  insularis  Casey  the  surface  is  more  shining  and  the  elytral 
punctures  are  finer,  the  pronotal  punctures  are  also  a  little  smaller  than 
in  latus.  The  characters  presented  by  this  phase  gradate  into  those  of 
latus. 

Infinitimus  Casey  has  not  been  recognized  in  the  material  before  me. 
The  habitat  is  uncertain  but  Casey  believes  the  type  came  from  the  Island 
of  Santa  Rosa.  A  large  series  of  specimens  collected  by  Mr.  Van  Duzee 
on  this  island  prove  to  be  a  true  Coniontis,  for  the  pronotal  base  is  almost 
truncate  from  angle  to  angle,  and  is  described  in  this  paper  as  santarosae. 
More  work  will  be  done  on  the  anatomy  of  these  forms. 

The  specimens  collected  by  Dr.  Van  Dyke  at  Los  Angeles  represent 
a  species  which  may  be  described  as  follows : 

Coniontides  vandykei,  new  species. — Form  elongate  oval,  about  twice 
as  long  as  wide,  moderately  convex ;  surface  moderately  shining.  Color 
black,  legs  and  tarsi,  antennae  and  mouth-parts  more  or  less  rufo-piceous. 

Head  quite  evenly  punctate,  punctures  well  defined  and  not  densely 
placed,  rather  more  numerous  anteriorly ;  epistomal  sinuation  shallow  and 
arcuately  reentrant.  Antennae  rather  short,  about  two-thirds  the  length 
of  the  pronotum ;  outer  joints  scarcely  wider  than  the  preceding  and 
moderately  compressed. 

Pronotum  rather  more  than  a  third  wider  at  base  than  long,  moder- 
ately and  evenly  convex  throughout ;  sides  moderately  arcuate,  more 
strongly  so  anteriorly,  apex  three-fifths  the  width  of  the  base,  lateral  bead 
rather  fine,  that  of  the  apex  very  fine,  base  not  beaded ;  disk  finely  and  not 
very  distinctly  punctate,  punctures  rather  evenly  but  not  closely  placed. 


EUSATTUS  DIFFICILIS  LEG.  213 

Elytra  evenly,  moderately  convex;  surface  very  finely  and  less  dis- 
tinctly punctate,  punctures  finer  than  those  of  the  pronotum,  becoming 
stronger  with  slight  rugulosity  on  the  apical  declivity. 

Prosternum  not  strongly  punctate,  punctures  rather  close,  sparse  on 
the  process,  which  is  not  margined  at  tip. 

Abdomen  finely,  rather  evenly  and  closely  punctate,  denser  on  the  fifth 
segment. 

Male.— Somewhat  narrower ;  pronotum  relatively  broader ;  abdomen 
less  convex. 

Female.— Slightly  broader;  pronotum  relatively  narrower;  abdomen 
more  convex. 

Measurements. — Length,  10  mm. ;  width,  5  mm. 

Type  locality.— Near  Los  Angeles,  California.  Collector,  Dr.  E.  C. 
Van  Dyke. 

Holotype,  female,  in  the  author's  collection ;  paratypes  in  the  collec- 
tion of  Dr.  Van  Dyke. 

Habitat. — Dr.  Van  Dyke  states  that  it  occurs  near  Los  Angeles,  at 
the  entrance  to  the  Griffith  National  Park. 

This  is  the  first  instance  of  finding  a  Coniontides  on  the  mainland. 
The  characteristics  of  vandykei  are  the  finer  punctuation  and  less  robust 
facies.  Seven  specimens  have  been  studied. 

Eusattus  difficilis  Lee. — A  large  series  of  this  species  is  before  me 
which  was  collected  at  San  Diego  and  Coronado,  in  1890  and  1891,  during 
the  months  of  October,  November,  April  and  June.  Part  of  the  series 
was  taken  by  Mr.  O.  N.  Sanford  and  Mr.  G.  W.  Dunn.  Casey  gives  San 
Diego  as  the  habitat  of  this  species.  My  series  gives  an  amazing  amount 
of  variation  in  the  characters  used  by  taxonomists  for  the  separation  of 
species.  Casey  considers  epipleural  characters  of  generic  value  and  states 
that  Horn  goes  so  far  as  to  give  such  trivial  characters  as  the  margination 
of  the  prosternal  lobe  precedence  over  the  epipleural  structure  (Proc. 
Washington  Acad.  Sciences,  10,  65,  1908).  I  am  discussing  here  intra- 
specific,  not  generic,  characters.  Margination  of  the  prosternal  process  is 
just  as  valuable  in  Eusattus  as  it  is  in  Coniontis,  and  it  is  of  no  positive 
value  in  either ;  but  is  helpful  in  a  limited  number  of  species.  Are  differ- 
ences only  to  be  considered  in  the  recognition  of  species,  and  must  a  pre- 
ponderance of  resemblances  be  overlooked  with  taxonomic  acumen? 
Study  a  series  of  Blapstinus,  Coniontis,  Eleodes,  Cryptadius,  Metoponium, 
Telabis,  Melanastus,  Coelocnemis,  etc.,  taken  from  a  limited  area  of  terri- 
tory and  what  are  ostensibly  the  same  species ;  compare  them  charac- 
ter for  character,  sex  for  sex.  Do  they  appear  to  have  been  cast  in  exactly 
the  same  mold  ?  Does  not  the  punctuation  vary,  also  the  margining  of  the 
prosternal  process,  body  form,  relative  length  of  the  appendages  in  the 


214      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

different  individuals  ?  So  it  is  with  my  series  of  Eusattus  difficilis,  taken 
on  the  same  small  peninsula,  about  the  same  bay,  same  climate  and  at 
different  seasons.  They  vary  in  surface  lustre,  from  opaque  to  somewhat 
shining,  prosternal  process  fully  margined  to  not  at  all,  elytra  with 
irregular  and  more  convex  longitudinal  lines  or  none  at  all,  sides  of  the 
pronotum  noticeably  declivo-explanate  to  scarcely  so,  pronotal  disk  with 
small  but  distinct  punctures  or  obsoletely  punctate,  prosternal  process 
narrow  or  broader, — all  these  variations  are  present  in  my  series  of  speci- 
mens, they  gradate  one  into  the  other.  So  far  the  diagnosis  is  clear. 

In  the  collection  of  the  California  Academy  of  Sciences  there  are  a 
few  specimens  of  Eusattus  taken  on  San  Martin  Island,  Lower  California, 
July  11,  1905,  by  F.  X.  Williams  while  on  the  Galapagos  Islands  Expe- 
dition. These  specimens  agree,  character  for  character,  with  certain  of 
the  specimens  collected  at  San  Diego.  The  individuals  of  the  small  series 
differ  among  themselves,  the  pronotum  is  distinctly  declivo-explanate 
laterally  in  one,  scarcely  so  in  another  and  to  an  intermediate  degree  in 
another.  The  most  surprising  thing  is  that  I  cannot  separate  them  by 
any  character  from  a  specimen  taken  by  Mr.  Frank  Daggett,  in  the  Kings 
River  Canyon,  California.  The  San  Martin  Island  specimens  are  smaller 
and  rather  more  oblong  than  the  average  difficilis,  but  agree  in  these 
characters  with  a  minority  of  specimens  from  San  Diego.  Does  geo- 
graphical position  make  the  San  Martin  Island  specimens  a  distinct  species 
from  those  taken  at  San  Diego?  Or,  am  I  right  in  my  conclusions  that 
they  are  all  of  one  species — namely,  difficilis  f 

Eusattus  vanduzeei,  new  species. — Form  oblong-oval,  strongly  con- 
vex, and  about  one-half  longer  than  wide.  Color  deep  black,  surface 
smooth  and  shining. 

Vertex  of  the  head  quite  impunctate,  elsewhere  rather  evenly  and  not 
closely  punctate,  punctures  rather  less  than  moderate  in  size,  densest  on 
the  epistoma,  the  latter  triangularly  sinuate  at  middle,  angles  rounded  into 
the  arcuate  sides,  which  are  feebly  or  not  sinuate  at  the  oblique  sutures, 
the  latter  may  or  may  not  be  evident ;  frontal  suture  more  or  less  distinct 
and  arcuate  between  the  'oblique  sutures ;  margin  of  the  epistomal  lobes 
slightly  reflexed. 

Pronotum  strongly  convex  from  side  to  side,  quite  obliquely  declivous 
laterally  and  rather  more  than  narrowly  explanate  at  the  sides ;  apex  and 
side  margins  finely  beaded,  base  not  beaded  but  sinuate  laterally,  median 
lobe  broad  and  feebly  arcuate;  basal  angles  obtuse  and  rather  narrowly 
rounded;  apical  angles  subobtuse;  disk  sparsely  punctate  laterally,  punc- 
tures fine  and  subobsolete  in  the  central  area,  coarser  laterally;  sides 
ciliate. 


EUSTATTUS  VANDUZEEI  215 

Elytra  with  very  fine  remotely  spaced  punctures ;  inflexed  sides  more 
distinctly  punctured  and  sometimes  subasperately  so,  each  puncture  with 
a  rather  long  hair ;  surface  sometimes  more  or  less  finely  wrinkled. 

Epipleura  finely  punctured,  each  puncture  with  a  hair  as  on  the 
inflexed  side  of  the  elytra ;  gradually  wider  from  apex  to  base. 

Body  beneath  pubescent,  each  puncture  bearing  a  hair.  Inflexed  sides 
of  the  pronotum  fimbriate  with  long  fulvous  hairs.  Pubescence  longest 
on  the  sterna  and  femora. 

Prosternum  asperato-punctate,  with  abundant  hairs;  process  mar- 
gined throughout;  surface  smooth  and  impunctate  between  and  behind 
the  coxae. 

Abdomen  very  finely  and  remotely  punctulate,  punctules  most 
abundant  laterally  and  on  the  fifth  segment,  hairs  short  and  scarcely 
evident  centrally. 

Anterior  tibiae  very  finely  serrulate  on  their  external  edge. 

Male. — Somewhat  narrower.  Pronotum  usually  as  wide  as  or 
slightly  wider  than  the  elytra.  Abdomen  less  convex. 

Female. — Broader  and  slightly  more  inflated.  Pronotum  as  wide  as 
the  elytra  or  slightly  narrower.  Abdomen  more  convex. 

Measurements. — Length  (Types),  12  mm.;  width,  7.0-7.5  mm. 

A  series  of  seventy-seven  specimens  studied.  The  individuals  are 
quite  uniform  as  regards  shape,  although  one  variant  is  subovate,  narrow- 
ing anteriorly,  with  the  pronotum  distinctly  narrower  than  the  elytra,  but 
continuing  the  side  lines  of  the  elytra.  Several  specimens  have  the  elytra 
distinctly  more  or  less  crinkled.  The  extremes  in  size  measure  as  follows : 
Length,  11-13  mm.;  width,  7-8  mm. 

Type  locality. — Prince  Island,  off  the  coast  of  Southern  California. 
Collected  by  E.  P.  Van  Duzee  on  May  20,  1919. 

Holotype,  male,  and  allotype,  female,  in  the  collection  of  the  Cali- 
fornia Academy  of  Sciences.  Paratypes  in  the  collection  of  the  Academy 
and  that  of  the  author. 

Habitat. — Prince  and  Santa  Rosa  Islands,  off  the  California  coast 
south  of  Santa  Barbara  County. 

This  fine  species  is  dedicated  to  its  discoverer.  There  has  been  some 
doubt  as  regards  its  relation  to  politus  Horn,  found  at  Santa  Barbara. 
While  vanduzcei  is  undoubtedly  related  to  politus,  it  can  not  be  identical 
if  the  published  descriptions  are  complete.  In  vanduzeei  the  sides  of  the 
pronotum  are  fimbriate,  the  inflexed  sides  of  the  elytra  and  epipleura 
clothed  with  fulvous  hairs,  and  the  under  surface  of  the  body  is  more  or 
less  pubescent  throughout.  The  sides  of  the  pronotum  are  somewhat 
explanate.  These  characters  are  not  mentioned  as  possessed  by  politus 
Horn.  In  vanduzeei  the  size  and  form  are  apparently  different. 


216      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Mr.  H.  C.  Fall  has  made  the  necessary  comparisons  at  Cambridge, 
and  found  two  examples  referred  to  politus.  "The  second  specimen  is  not 
politus  at  all.  The  first  bears  the  name  label  in  Horn's  handwriting  and 
is  evidently  one  of  his  type  series.  It  is  rather  old  and  worn,  hence  the 
setae  are  not  well  preserved.  They  are  present,  however,  on  the  inflexed 
sides  of  the  thorax,  but  appear  to  be  practically  absent  from  the  epipleura, 
nor  is  there  any  evidence  of  setigerous  punctures  to  show  where  they  may 
have  been,  except  possibly  two  or  three  very  small  ones."  He  further 
states  that  "the  punctuation  of  the  upper  surface  is  distinct  enough 
though  sparse  and  fine.  Your  specimens  are  distinctly  smoother." 

Mr.  Van  Duzee  states  that  the  specimens  were  "taken  on  the  steep 
southern  slope  of  Prince  Island  at  the  entrance  to  the  harbor  of  San 
Miguel  Island.  They  were  found  beneath  prostrate  weeds  more  or  less 
imbedded  in  light  sandy  soil  brought  down  by  the  small  surface  washes. 
In  the  right  kind  of  soil  they  were  not  uncommon,  an  hour's  work  yielding 
about  seventy  specimens. 

The  mentum  in  vanduzeei  is  transverse,  with  the  apex  broadly  sinuate, 
sinuation  nearly  semicircular  at  middle  third,  thence  oblique  and  feebly 
arcuate  to  the  tips  of  the  lobes,  the  latter  are  obtusely  rounded ;  sides  dis- 
tinctly arcuate;  surface  nearly  impunctate  centrally,  broadly  impressed 
laterally  on  the  lobes  and  punctate,  impression  deep  at  base ;  each  puncture 
bearing  a  yellow  seta ;  lateral  bead  thick. 

In  Eusattus  difficilis  the  surface  of  the  mentum  is  feebly  convex, 
nearly  impunctate,  with  only  a  few  scattered  punctures ;  otherwise  as  in 
vanduzeei.  The  mentum  is  still  smoother  in  Eusattus  reticulatus  Say. 

Female  genital  segment. — Each  valve  consists  of  a  dorsal,  a  lateral 
plate  terminating  in  the  apex,  and  a  ventral  plate.  Lateral  plate  strongly 
chitinized,  dilated  at  base  but  continuous  with  the  less  chitinized  dorsal 
and  ventral  plates ;  surface  with  very  sparse  punctures,  each  with  a  long 
hair;  apex  slightly  recurved,  flattened  dorso-internally,  feebly  convex  on 
the  ventro-lateral  surface ;  the  rather  elongate  oval  fossa  bears  a  pencil  of 
about  eight  long  hairs  situated  near  base  of  the  apex ;  fossa  membranous 
at  bottom ;  tip  of  apex  strongly  chitinized  and  narrowly  rounded.  Dorsal 
plate  pale  in  color,  not  moderately  chitinized,  internal  border  apparently 
arcuate,  reaching  to  a  point  about  midway  between  the  fossa  and  tip  of 
apex ;  surface  coarsely  and  rather  closely  punctate,  each  puncture  bearing 
a  long  hair.  Ventral  plate  narrower  and  elongate,  pale  in  color  and  not 
strongly  chitinized,  reaching  to  a  point  opposite  the  tip  of  the  ventral 
plate ;  surface  densely  and  coarsely  punctate,  each  puncture  bearing  a  long 
hair.  Dorsal  and  ventral  plates  blend  with  the  lateral  plate.  Described 
from  the  dried  specimen. 


ELEODES  ROTUNDIPENNIS  217 

Eleodes  rotimdipennis  Lee.— Recent  studies  demand  that  rotundi- 
pennis  be  raised  from  the  grade  of  variety  to  that  of  a  valid  species,  with 
the  following  brief  diagnosis :  Pronotal  sides  evenly  and  broadly  rounded 
(as  in  parvicollis),  basal  constriction  short  and  the  angles  rectangular. 
Otherwise  as  in  cor  data  Esch.  It  should  follow  scabrosa  Lee.  and  precede 
cordata  in  our  lists.  For  further  remarks  see  the  author's  Monograph  of 
the  Eleodiini  (Bull.  63,  U.  S.  Nat.  Mus.). 

Habitat. — Oregon  (Koebele)  ;  British  Columbia  (Victoria  and  North 
Bend. — Hubbard  and  Schwarz)  ;  State  of  Washington  (Easton  and 
Olympia). 

Eleodes  rotundipennis  versatilis,  new  variety. — Form  elongate  sub- 
oblong,  not  strongly  convex  and  a  little  more  than  twice  as  long  as  wide. 
Color  piceous  to  deep  black,  more  or  less  shining.  Pronotum  rather 
strongly  and  moderately  coarsely  punctate,  punctures  irregularly  dis- 
tributed, leaving  small  smooth  areas ;  disk  flat  between  punctures.  Elytra 
muricato-tuberculate,  scarcely  rugose. 

Head  feebly  convex,  rather  closely  punctate,  punctures  rather  less 
than  moderate  and  more  or  less  evenly  distributed;  impressions  broad, 
shallow  and  within  the  antennal  prominences.  Antennae  rather  long, 
slightly  incrassate ;  ninth  and  tenth  joints  slightly  transverse,  eleventh  sub- 
trapezoidal. 

Pronotum  rather  transversely  oblong,  moderately  and  evenly  convex, 
widest  at  middle ;  disk  quite  evenly  convex,  punctures  more  or  less  dis- 
crete, moderate  in  size  and  distinct,  irregular  in  distribution,  leaving  small 
glabrous  areas,  slightly  denser  laterally  and  feebly  subasperate,  intervals 
flat;  sides  rather  evenly  and  broadly  arcuate  in  anterior  five-sixths,  sud- 
denly constricted  at  base,  where  the  sides  are  straight  and  parallel,  mar- 
ginal bead  fine;  apex  feebly  emarginate  in  a  circular  arc;  apical  angles 
obtuse  and  not  in  the  least  prominent ;  base  truncate,  equal  to  the  apex ; 
basal  angles  rectangular. 

Propleura  opaque,  sparsely  and  muricately  punctate. 

Elytra  distinctly  oblong-oval,  moderately  convex,  widest  in  the  middle 
third;  base  truncate,  scarcely  wider  than  the  pronotum,  humeri  distinct 
and  obtusely  rounded;  sides  evenly  and  not  strongb/  arcuate,  obliquely 
converging  to  apex  in  apical  third,  apex  obtusely  rounded ;  disk  arcuately 
declivous  at  apex,  evenly  and  not  very  closely  muricately  punctate  but  not 
strongly  so,  punctures  becoming  quite  tuberculate  laterally  and  on  apex, 
tubercles  shining  at  summit. 

Abdomen  rather  strongly  and  densely  punctate,  except  on  the  fourth 
and  fifth  segments,  which  are  finely  punctate. 

Legs  moderate  in  length ;  anterior  femora  mutic  as  usual  in  the  sub- 
genus.  Anterior  and  middle  tarsi  dissimilar  in  the  sexes. 


218      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Male. — Somewhat  narrow  and  parallel,  antennae  extending  to  or  just 
beyond  the  pronotal  base.  Abdomen  moderately  convex,  slightly  oblique 
to  the  sterna,  somewhat  flattened  along  the  middle.  First  three  joints  of 
anterior  tarsi  with  small  subacute  tufts  of  yellowish  pubescence  at  tips 
beneath,  that  of  the  third  joint  smaller  than  the  others ;  first  two  joints 
of  the  middle  tarsi  with  similar  and  smaller  tufts. 

Female. — More  robust.  Abdomen  more  convex.  First  joint  of  the 
anterior  tarsi  slightly  thickened  at  apex  beneath,  plantar  groove  more  or 
less  distinct  throughout. 

Measurements. — Length,  14.5  mm. ;  width,  6  mm. 

Holotype,  male,  in  the  collection  of  the  California  Academy  of 
Sciences.  Paratype  in  the  academy's  and  the  author's  collection. 

Type  locality. — Colestin,  Jackson  County,  Oregon.  Collected  July 
30,  1918,  by  E.  P.  Van  Duzee ;  one  specimen  taken  at  Easton,  Washington. 

Versatilis  was  one  of  the  puzzling  forms  studied  at  tht  time  the 
Monograph  of  the  Eleodiini  was  written,  and  no  decision  was  reached. 
Two  closely  related  forms  are  still  doubtful,  these  having  been  collected 
in  central  and  southern  California. 

Versatilis  has  the  pronotum  of  parvicollis  and  the  sculpturing  of 
cordata. 

Eleodes  dentipes  marinae,  new  variety. — Form  moderately  robust, 
oval  to  ovate  and  almost  subfusiform,  convex  and  very  smooth,  punctua- 
tion fine  and  distinct ;  punctures  subequal  throughout. 

Head  feebly  convex,  impressions  obsolete,  punctures  rather  irregu- 
larly distributed.  Antennae  rather  short,  not  reaching  to  base  of  the 
pronotum,  gradually  incrassate  to  tip,  ninth  and  tenth  joints  transverse, 
eleventh  obliquely  truncate  at  apex. 

Pronotum  widest  at  middle,  about  two-sevenths  wider  than  long ;  disk 
rather  evenly  and  strongly  convex,  rather  strongly  declivous  at  the  apical 
angles,  finely,  distinctly,  not  closely  punctate  throughout ;  punctures  equal 
in  size  and  distinct  to  the  margin,  lateral  bead  fine;  sides  evenly  and 
moderately  strongly  arcuate  in  anterior  two-thirds,  thence  straight  and 
convergent  almost  to  the  basal  angles,  which  are  slightly  prominent 
laterally,  making  the  margin  feebly  sinuate  just  in  front  of  them;  apex 
broadly  and  feebly  emarginate,  feebly  arcuate  in  middle  third ;  apical 
angles  short,  rather  stout,  somewhat  prominent,  sometimes  more  or  less 
everted ;  base  feebly  arcuate. 

Propleura  smooth,  feebly,  sparsely  punctate  and  more  or  less  rugu- 
lose. 

Elytra  oval,  widest  at  middle,  smooth,  quite  evenly  convex  from  side 
to  side ;  punctuation  fine,  punctures  evenly  distributed,  not  close  and  equal 


ELEODES  LATICOLLIS  APPRIMA  219 

in  size,  diffuse  or  with  striae  of  punctures  more  or  less  evident ;  base 
feebly  emarginate ;  disk  rather  gradually  and  arcuately  declivous  apically, 
the  apex  obtusely  rounded ;  sides  arcuate  throughout,  converging  to  apex. 

Abdomen  finely  and  sparsely  punctured. 

Legs  moderate  in  length. 

Male. — Narrower,  elongate,  subfusiform  oval.  Elytra  gradually  nar- 
rowed posteriorly,  rather  gradually  and  arcuately  declivous  apically ;  apex 
obtusely  rounded,  sometimes  feebly  divergent  at  tip.  Abdomen  moder- 
ately convex,  first  two  segments  scarcely  impressed  or  flattened.  Anterior 
femora  with  a  slender  and  well  developed  tooth.  Posterior  femora 
slightly  arcuate,  gradually  widened  toward  apex. 

Female. — Rather  robust.  Elytra  more  or  less  broadly  oval,  rather 
strongly  although  gradually  declivous  posteriorly ;  apex  rather  more 
broadly  rounded.  Abdomen  rather  moderately  and  evenly  convex. 
Anterior  femora  with  an  obtuse  tooth,  posterior  quite  straight. 

Measurements. — Length   (Types),  22-24  mm.;  width,  8.2-9.5  mm. 

Holotype,  female,  and  allotype,  male,  in  the  collection  of  the  Cali- 
fornia Academy  of  Sciences.  Paratypes  in  the  Academy's  and  the 
author's  collection. 

Type  locality. — Fairfax,  Marin  County,  California,  March  9,  1919; 
E.  P.  Van  Duzee,  collector. 

Habitat. — As  far  as  known,  found  only  in  Marin  County,  California. 

A  moderate  series  has  been  studied.  Marinae  caused  considerable 
trouble  until  Mr.  Van  Duzee  took  a  colony  which  adhered  to  a  common 
type  form.  Marinae  should  precede  confinis  Blais.  in  our  lists.  In  con- 
finis  the  pronotal  sides  posteriorly  are  straighter  to  the  angles  than  in 
marinae;  there  is  a  shallow  and  broad  sinuation  in  front  of  the  basal 
angles,  the  latter  being  somewhat  prominent  laterally  in  marinae  while  in 
confinis  the  angles  are  not  at  all  prominent;  in  confinis  the  punctuation 
is  fine  and  indistinct ;  in  marinae  it  is  less  fine,  distinct  and  well  defined. 

Eleodes  laticollis  apprima,  new  subspecies. — Since  the  publication  of 
the  Monograph  of  the  Eleodiini,  a  large  series  of  specimens  of  what  was 
then  referred  to  Eleodes  laticollis  forma  insularis,  has  been  studied  and  the 
resulting  conclusion  is  that  it  is  a  valid  subspecies  and  should  be  raised  to 
that  grade,  as  indicated  above.  The  material  at  hand  includes  collections 
made  on  San  Nicolas,  Prince,  San  Clemente,  Santa  Barbara,  Anacapa  and 
San  Miguel  Islands.  Material  ample  enough  to  force  a  change  of 
opinion. 

In  apprima  the  pronotum  is  as  a  rule  a  little  longer  when  compared 
with  the  width,  notably  shining,  and  the  form  is  more  elongately  robust. 
The  individuals  vary  in  every  way  as  they  do  in  laticollis. 


220      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Measurements. — Length  (Types),  18-20.5  mm.;  width,  10-12.5  mm. 

Holotype,  female,  and  allotype,  male,  in  the  collection  of  the  Cali- 
fornia Academy  of  Sciences ;  paratypes  in  the  Academy's  and  the  author's 
collection. 

Type  locality.— San  Nicolas  Island,  March  28,  1918;  J.  R.  Slevin, 
collector. 

The  characters  are  quite  similar  in  laticollis  and  apprima ;  the  former 
has  a  relatively  more  transverse  pronotum  and  is  usually  dull  in  lustre. 

Eleodes  sanmartinensis,  new  species. — Form  elongate,  oblong-oval  to 
suboblong-oval.  Color  black  and  more  or  less  shining;  surface  smooth. 

Head  about  as  long  as  wide,  equal  to  one-half  the  width  of  the 
pronotum ;  front  slightly  convex,  punctures  small  and  irregular  in  dis- 
tribution, sparse  centrally  and  denser  laterally.  Antennae  extend  slightly 
beyond  the  base  of  the  pronotum,  feebly  incrassate  apically,  last  three 
joints  scarcely  wider  and  more  or  less  globular. 

Pronotum  about  a  fifth  wider  than  long;  sides  evenly  and  broadly 
arcuate  from  apex  to  base,  marginal  bead  more  or  less  moderate  and 
slightly  reflexed ;  apex  more  or  less  truncate  between  the  rather  small  and 
more  or  less  everted  apical  angles,  finely  beaded,  bead  more  or  less  obsolete 
at  middle ;  base  truncate,  rather  finely  and  evenly  beaded ;  disk  evenly  and 
moderately  convex,  rather  narrowly  impressed  along  the  sides,  punctua- 
tion fine  and  more  or  less  obsolete,  except  laterally  where  the  punctures 
are  more  distinct  and  slightly  punctato-rugulose  on  the  impressed  area, 
surface  smooth ;  basal  angles  obtuse  and  more  or  less  distinct. 

Propleura  smooth  and  obsoletely  rugose. 

Elytra  more  or  less  oval,  humeri  obsolete;  sides. evenly  arcuate,  in 
apical  fourth  evenly  converging  to  the  obtuse  apex ;  base  transverse ;  disk 
smooth  and  more  or  less  indistinctly  sculptured ;  punctures  small  and  sub- 
equal,  not  in  the  least  asperate,  apparently  more  or  less  serial  in  arrange- 
ment, double  series  sometimes  present. 

Parapleura  more  or  less  coarsely  punctured. 

Sterna  not  strongly  sculptured. 

Abdomen  smooth,  finely  and  sparsely  punctate,  punctures  denser  on 
the  fifth  segment,  more  or  less  rugulose. 

Legs  long,  femora  heavy  and  markedly  compressed,  anterior  femora 
dentate;  tibiae  densely  punctato-muricate.  Tarsi  apparently  similar  in 
the  sexes. 

Male. — More  or  less  parallel  and  elongate  oblong-oval.  Elytra  not 
wider  than  the  pronotum,  more  gradually  declivous  posteriorly.  Femora 
notably  stout,  long  and  compressed,  the  anterior  distinctly  swollen,  tooth 
at  outer  fourth,  subcylindric,  everted  and  backwardly  curved,  resulting 


ELEODES  SANMARTINENSIS   MOESTA  221 

apical  sinus  more  or  less  arcuately  narrowed;  metatibiae  quite  straight. 
Abdomen  notably  flattened  and  moderately  oblique  to  the  sterna. 

Measurements. — Length,  22.9  mm. ;  width,  9.5  mm. 

Female. — Suboblong-oval,  elytra  moderately  inflated,  distinctly  wider 
than  the  pronotum,  more  abruptly  declivous  posteriorly.  Femora  less 
stout,  tooth  of  pro  femora  less  stout,  sinus  more  open.  Abdomen  moder- 
ately convex  and  on  the  same  plane  with  the  sterna. 

Measurements. — Length,  26  mm.;  width,  11  mm. 

Holotype,  female,  in  the  collection  of  the  California  Academy  of 
Sciences.  Allotype,  male,  in  the  author's  collection. 

Type  locality  and  habitat. — San  Martin  Island,  off  the  coast  of 
northern  Lower  California,  July  11,  1905;  F.  X.  Williams  collector. 

Sanmartinensis  differs  from  laticollis  subspecies  apprima  in  the 
broadly  rounded  sides  of  the  pronotum,  without  basal  constriction.  An 
unusual  character  in  the  dentipes  and  laticollis  sections  of  the  genus.  The 
general  form  is  that  of  a  male  grandicollis  Mann. 

Eleodes  sanmartinensis  moesta,  new  variety. — Form  elongate  ovate, 
surface  lustre  dull,  elytra  sparsely  and  muricately  sculptured. 

Sides  of  head  in  front  of  the  antennae  rather  rapidly  converging, 
frons  rather  flat,  sparsely,  finely  and  distinctly  punctate.  Antennae  rather 
short  and  slender,  ninth  and  tenth  joints  transversely  oval,  eleventh  trun- 
cate oval. 

Pronotum  a  little  wider  than  long ;  sides  broadly  arcuate,  somewhat 
more  strongly  so  in  anterior  half,  thence  somewhat  straight  to  the  basal 
angles ;  punctuation  of  the  disk  fine,  sparse  and  distinct. 

Elytra  suboval,  humeri  obtuse  but  distinct;  disk  feebly  flattened  in 
the  central  area,  punctuation  consisting  of  small,  distinct  granulate  punc- 
tures ;  granules  shining  on  summit ;  each  puncture  with  a  distinct  black 
semi-erect  seta,  punctures  serially  arranged  in  the  central  part  of  the 
disk  with  an  interstitial  and  more  widely  spaced  row ;  punctures  fine,  be- 
coming rather  coarse  and  asperate  on  the  sides  and  apical  declivity ;  sides 
rather  broadly  arcuate,  somewhat  oblique  in  apical  fourth  to  apex,  the 
latter  obtuse. 

Legs  densely  sculptured  and  comparatively  short. 

Otherwise  as  in  sanmartinensis. 

Female. — Abdomen  convex ;  femora  normal  in  thickness. 

Measurements.— Length,  21  mm. ;  width,  9.5  mm. 

Holotype,  female,  in  the  collection  of  the  California  Academy  of 
Sciences. 

Type  locality  and  habitat. — San  Martin  Island,  off  the  coast  of 
northern  Lower  "California.  Collected  July  11,  1905;  F.  X.  Williams, 
collector. 


222      NEW  SPECIES   MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Moesta  bears  the  same  relation  to  sanmartinensis  that  var.  minor  does 
to  laticollis  Lee.  There  is  a  slight  indication  of  an  obtuse  salient  at  apical 
fourth  of  the  mesofemora.  The  present  race  is  based  on  analogy  that  has 
been  observed  in  series  of  many  species.  Moesta  differs  so  radically  in 
sculpturing  from  the  type  species  that  it  can  not  be  considered  as  hetero- 
typical  and  therefore  a  form. 

Eleodes  mutilata,  new  species. — Form  elongate  ovate  to  fusiform 
ovate,  convex.  Color  black,  surface  smooth,  femora  edentate;  legs 
slender  and  moderate  in  length. 

Head  moderately  small,  rather  short,  front  moderately  convex, 
epistomal  suture  fine  and  more  or  less  impressed;  surface  smooth,  very 
finely  and  sparsely  punctate.  Antennae  long  and  slender,  reaching  about 
two  joints  beyond  the  pronotal  base ;  joints  four  to  eight  inclusive  elongate 
and  subequal,  ninth  and  tenth  triangulo-oval,  eleventh  obovate,  pointed  at 
apex  and  about  as  long  as  wide,  last  three  joints  scarcely  wider  to 
slightly  so. 

Pronotum  quadrate,  a  little  wider  than  long;  apex  broadly  and  not 
strongly  emarginate  between  the  anteriorly  prominent  angles,  the  latter 
subacute  and  rather  broadly  subdentiform ;  base  broadly  and  feebly 
arcuate,  apex  and  base  obsoletely  beaded ;  sides  broadly  and  not  strongly 
arcuate,  very  slightly  sinuate  behind  and  before  the  apical  and  basal 
angles,  the  latter  subobtuse  and  not  at  all  prominent,  marginal  bead  very 
fine,  almost  subobsolete ;  disk  rather  strongly,  evenly  convex  and  impunc- 
tate,  or  with  fine,  sparse  and  subobsolete  punctures. 

Propleura  sparsely  and  obsoletely  punctate  and  rugose. 

Elytra  oval  to  subfusiform-oval,  rather  strongly  convex,  sometimes 
slightly  flattened  on  the  dorsum ;  sides  evenly  arcuate,  slightly  more 
oblique  in  apical  third,  apex  obtusely  rounded ;  disk  smooth,  obsoletely 
sculptured,  punctures  small  and  subequal  in  size  throughout  and  arranged 
in  closely  placed  series ;  base  feebly  and  broadly  emarginate,  subequal  to 
the  pronotal  base ;  humeri  minutely  dentiform. 

Parapleura  rather  distinctly  and  densely  punctate. 

Sterna  more  or  less  obsoletely  sculptured. 

Abdomen  smooth,  apical  half  of  fifth  segment  finely  and  densely 
punctured ;  surface  more  or  less  obsoletely  and  irregularly  rugose. 

Legs  rather  feebly  sculptured,  except  the  distal  half  of  the  tibiae, 
which  are  densely  muricato-punctate.  Tarsi  dissimilar  in  the  sexes. 
Femora  mutic. 

Male. — Rather  more  elongate,  less  convex  and  a  little  narrower. 
Abdomen  slightly  oblique  to  the  sterna,  feebly  flattened.  First  two  joints 
of  the  anterior  tarsi  clothed  beneath  with  pads  of  rather  coarse  golden 


ELEODES  MUTILATA  223 

brown  setiform  hairs,  which  are  graduated  from  base  to  apex;  similar 
setiform  hairs  form  apico-marginal  tufts  on  the  other  joints,  divided  at 
middle,  and  leave  the  plantar  grooves  open ;  on  the  middle  tarsi  the 
plantar  grooves  are  open,  the  joints  are  fringed  beneath  with  golden 
brown  setiform  hairs. 

Female. — Stouter,  body  rather  more  inflated  and  broader. 
Abdomen  horizontal,  rather  evenly  and  not  strongly  convex.  Plantar 
grooves  open  on  all  of  the  tarsi;  joints  with  apico-marginal  tufts  of 
golden  brown  setiform  hairs. 

Measurements. — Length  (Types),  28-26  mm.;  width,  11  mm. 

Holotype,  male,  in  the  collection  of  the  California  Academy  of 
Sciences ;  paratypes  in  the  Academy's  and  the  author's  collection.  Three 
specimens  studied. 

Type  locality  and  habitat. — Sierra  Laguna,  Lower  California;  col- 
lected by  J.  R.  Slevin  on  August  15,  1919. 

The  form  and  habitus  are  exactly  alike  in  mutilata  and  lucae  Lee.  It 
has  the  appearance  of  being  an  edentate  and  smooth  lucae.  The  anterior 
femora  are  entirely  mutic.  As  nothing  is  known  regarding  the  status 
of  similar  forms  it  is  best  to  consider  it  a  species  rather  than  a  race  of 
lucae. 

The  following  are  the  genital  characters : 

Male. — Edeagophore  oblong-ovate,  moderate  in  size,  about  two  and 
a  half  times  longer  than  wide  and  feebly  arched. 

Basale  not  strongly  convex,  less  so  on  the  dorsum,  with  an  impressed 
median  groove ;  sides  parallel,  less  than  moderately  and  broadly  arcuate, 
apex  bilobed,  median  emargination  arcuately  rounded  at  bottom. 

Apicale  triangular,  apex  short  and  rather  obtusely  rounded ;  sides 
broadly  arcuate,  but  briefly  sinuate  at  sides  of  apex ;  surface  convex 
with  an  impressed  median  groove  extending  from  apex  to  base ;  base 
sinuate  laterally,  adapted  to  the  apical  lobes  of  the  basale,  obtusely  lobed 
at  middle. 

Female. — Genital  segment  subtriangulo-parabolic,  moderate  in  size 
and  setose. 

Valvula. — Dorsal  plate  oblong,  rather  narrow,  sides  feebly  arcuate, 
with  apex  rounded;  surface  feebly  concave,  finely  and  very  sparsely 
punctate,  each  puncture  with  a  very  small  seta.  Apex  short,  stout, 
rather  broadly  rounded  at  tip,  somewhat  divergent  with  that  of  the 
opposite  valve.  Appendage  situated  at  and  almost  beneath  the  apical 
margin  of  the  dorsal  plate,  very  small  and  not  easily  seen  within  the 
fossa.  Fossa  moderate  in  size,  adjacent  surface  of  apex  with  many 
short  hairs.  Superior  pudendal  membrane  attaining  the  middle  of  the 


224      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

dorsal    plate,   broadly   and   rather   parabolically   exposed,    long-itudinally 
rugulose. 

Valvular  membranes  visible  between  the  apices  and  finely  setose. 
Ventro-lateral  surfaces  evenly  convex,  not  noticeably  swollen ;  sub- 
marginal  groove  moderate  and  continuous  with  the  concavity  on  the 
under  surface  of  the  apex ;  surface  with  very  fine  and  very  sparsely 
placed  punctures  and  setae.  Internal  margins  of  the  valves  contiguous 
apically  and  briefly  so  at  base.  Genital  fissure  ample  and  fusiform, 
exposing  the  inferior  pudendal  membrane. 

Eleodes  corvina,  new  species. — Form  elongate  subovate,  about  twice 
as  long  as  wide,  moderately  convex.  Color  deep  black,  rather  shining, 
head  and  pronotum  somewhat  duller. 

Head  short,  wider  than  long,  feebly  convex,  frontal  suture  more  or 
less  distinct  and  moderately  impressed;  surface  closely  and  evenly 
punctured,  punctures  almost  coarse  and  more  or  less  coalescent. 
Antennae  moderate  in  length,  reaching  to  the  base  of  the  pronotum, 
gradually  and  feebly  incrassate;  third  joint  equal  to  the  fourth  and  fifth 
taken  together ;  fifth,  sixth  and  seventh  subequal  in  length  and  very  little 
longer  than  wide;  eighth  subtriangular,  ninth  triangulo-oval,  tenth 
transversely  oval ;  eleventh  short  ovate,  about  as  long  as  wide,  rather 
obliquely  truncate  at  tip. 

Pronotum  moderately  transverse,  widest  at  middle,  apex  almost 
truncate,  apical  angles  distinct,  not  in  the  least  prominent  and  obtuse ; 
sides  rather  strongly  and  broadly  arcuate,  almost  straight  posteriorly 
and  convergent  to  the  base,  marginal  bead  fine;  base  slightly  arcuate  to 
subtruncate,  obsoletely  beaded ;  basal  angles  obtuse  and  deflexed  with 
the  discal  surface ;  disk  rather  strongly  convex,  most  strongly  so  laterally 
and  at  the  basal  angles,  where  it  is  quite  declivous ;  densely,  somewhat 
irregularly  punctate,  punctures  a  little  larger  than  those  of  the  head, 
slightly  coarse,  rather  smaller  and  sparsely  placed  centrally,  becoming 
more  or  less  crowded  and  coalescent  laterally. 

Propleura  full  and  convex,  more  or  less  but  not  strongly  muricately 
punctato-rugose. 

Elytra  elongately  ovate  to  oval,  moderately  convex,  obliquely  and 
rather  rapidly  declivous  posteriorly,  apex  rather  narrowly  rounded; 
humeri  distinct,  not  in  the  least  prominent;  base  feebly  emarginate  and 
a  little  wider  than  the  contiguous  pronotal  base;  disk  distinctly  and 
strongly  sculptured,  rather  coarsely  punctato-rugose  centrally  along  the 
suture,  becoming  rugoso-tuberculate  laterally  and  on  the  apical 
declivity. 

Parapleura  and  sterna  rather  densely  and  not  coarsely  punctate. 

Abdomen  rather  densely  punctate  and  more  or  less  rugose. 


ELEODES   ACUTANGULA  225 

Legs  somewhat  short  and  moderately  stout,  rather  densely  sculp- 
tured. Tarsi  similar  in  the  sexes,  plantar  grooves  open  throughout. 

Male. — Narrower  and  more  elongate  ovate,  somewhat  less  convex. 
Abdomen  slightly  oblique  to  the  sterna  and  noticeably  flattened ;  tibial 
spurs  short. 

Female. — Broader,  more  elongate  oval,  elytra  more  convex ; 
abdomen  horizontal  and  more  convex. 

Measurements. — Length  (Types),  14-15  mm.;  width,  5-6  mm. 

Holotype,  male,  and  allotype,  female,  in  my  own  collection.  Para- 
types  in  that  of  Mr.  Ralph  Hopping. 

Type  locality. — Dry  Lake,  Klamath  National  Forest.  Collected  on 
September  17,  1912,  by  Mr.  Ralph  Hopping. 

The  elytral  sculpturing  in  corvina  is  almost  like  that  observed  in 
Eleodes  cuneaticollis  Casey,  but  more  coarsely  punctate  centrally  and 
more  distinctly  tuberculate  laterally.  Papillosa  Blais.,  a  closely  related 
congeneric  species,  is  larger  and  more  elongate,  the  elytral  tubercles  are 
rounded,  distinct,  shining  at  their  summits  and  not  intermixed  with 
rugae ;  in  granulata  Lee.  the  surface  is  more  opaque  and  the  elytral 
tubercles  are  rather  elongate,  while  in  the  variety  obtusa  Lee.  the  sculp- 
turing is  more  eroded  and  dull,  with  the  body  form  more  like  that  of 
corvina,  but  in  the  latter  the  elytral  sculpturing  is  distinctly  punctato- 
tuberculato-rugose.  The  sculpturing  of  vandykei  Blais.  is  much  less 
developed  and  more  of  the  muricato-punctate  type,  while  in  its  race 
parvula  Blais.  the  size  is  smaller,  the  pronotum  more  quadrate,  and  the 
elytral  tubercles  are  evident,  variable  in  number  and  more  or  less 
muricato-tuberculate. 

Eleodes  acutangula,  new  species. — Form  oblong-oval  and  moder- 
ately convex  as  in  scabricula  Lee.  Color  black  and  quite  opaque. 

Head  rather  less  than  one-half  as  wide  as  the  pronotum,  feebly 
convex  with  broad  and  very  shallow  impressions  within  the  antennal 
convexities ;  rather  coarsely  and  irregularly  punctate.  Antennae  about 
attaining  the  pronotal  base,  outer  joints  feebly  compressed  and  slightly 
wider. 

Pronotum  distinctly  opaque,  widest  at  middle  and  transverse ;  apex 
broadly,  evenly  and  moderately  emarginate  in  circular  arc ;  sides  quite 
evenly  and  broadly  arcuate,  straighter  posteriorly  to  base,  margin  more 
or  less  feebly  reflexed ;  apical  angles  quite  prominent  anteriorly  and 
rather  acute ;  basal  angles  obtuse,  distinct  and  slightly  prominent  laterally 
on  account  of  the  basal  bead. 

Elytra  oval,  opaque,  sculptured  as  in  consobrina  Lee.,  but  somewhat 
coarser,  that  is,  rather  coarsely  punctate  centrally  and  granulate- 


226      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

muricate  laterally  and  on  the  apical  declivity ;  humeri  rather  distinct ;  disk 
moderately  convex  to  somewhat  flattened  at  times. 

Otherwise  as  in  scabricula. 

Male. — Elongate  oval  and  narrower. 

Female. — More  robust. 

Measurements. — Length  (Types),  20-21  mm.;  width,  9-10  mm. 

Holotype,  male,  and  allotype,  female,  in  my  own  collection.  Para- 
types  in  that  of  Mr.  Ralph  Hopping. 

Type  locality. — Cannel  Meadow,  Kern  County,  California ;  col- 
lected June  11,  1913,  by  Mr.  Ralph  Hopping. 

Habitat. — Cannel  Meadow  and  Breckenridge  Mountain,  Kern 
County,  California.  A  paratype  from  the  latter  locality  was  taken  at  an 
altitude  of  6000  feet.  Number  of  specimens  studied,  nine. 

Acutangula  differs  from  scabricula  in  its  distinct  and  quite  strong 
apical  pronotal  angles,  coarser  sculpturing  and  more  opaque  lustre.  In 
scabricula  the  apical  angles  are  distinctly  obtuse  and  rather  narrowly 
rounded,  basal  angles  obtuse  and  not  in  the  least  prominent.  In 
acutangula  the  basal  angles  are  distinct  and  obtuse,  and  rendered  slightly 
prominent  by  the  basal  bead  alone. 

A  single  specimen  taken  by  Mr.  J.  R.  Slevin,  at  Glenbrook,  Nevada, 
on  August  25,  1913,  is  referred  to  this  species. 

Tenebrio  tenebroides  Beauv. — This  phase  or  race  of  T.  picipes 
Herbst.,  according  to  the  Leng  Catalogue  of  the  Coleoptera  of  North 
America,  has  a  wide  range  of  distribution,  being  particularly  mentioned 
as  occurring  in  Southern  California,  Newfoundland  and  Florida.  In  all 
probability  it  is  only  one  of  the  modifications  of  the  cosmopolitan  picipes. 

Some  thirty  years  ago  I  took  a  single  specimen  from  beneath  the  bark 
of  a  tree  on  top  of  the  Bolcan  Mountain,  near  Julian,  San  Diego  County, 
California.  This  region  was  isolated  as  far  as  commerce  was  concerned, 
but  not  far  distant  from  the  Santa  Isabel  Indian  Reservation.  At  any 
rate  the  specimen  remained  a  unique  in  my  collection  until  quite  recently, 
when  Mr.  F.  W.  Nunenmacher  found  another  in  Shasta  County,  and 
meanwhile  Mr.  Ralph  Hopping  collected  another  at  Fort  Tejon,  California. 
All  these  localities  were  away  from  commercial  warehouses.  These 
specimens  differ  from  the  eastern  examples  in  being  more  distinctly 
punctured  on  the  upper  surface.  In  the  more  abundant  specimens  of 
picipes  Herbst.  the  punctuation  of  the  head,  pronotum  and  elytral  intervals 
is  scarcely  distinct.  In  both  forms  the  punctures  of  the  elytral  striae  are 
distinct.  The  manner  of  dispersion  of  cosmopolitan  species  is  an  inter- 
esting problem. 

Helops  simulator,  new  species. — Form  elongate-ovate,  alate;  wings 
about  as  long  as  the  elytra ;  densely  punctate  above.  Color  black. 


TENEBRIO  TENEBROIDES  BEAUV.  227 

Head  about  as  long  as  wide,  transversely  impressed  on  the  epistoma ; 
front  scarcely  convex,  densely  punctate,  punctures  moderate  in  size. 
Antennae  moderately  long  and  slender,  outer  joints  compressed. 

Pronotum  subquadrate,  slightly  wider  than  long;  apex  truncate;  base 
subtruncate,  very  feebly  sinuate  laterally;  sides  moderately  arcuate  in 
anterior  two-thirds,  thence  slightly  convergent  to  base  and  more  or  less 
sinuate  to  parallel  before  the  angles ;  disk  moderately  convex,  declivous  at 
the  apical  angles  and  slightly  flattened  about  the  basal  angles,  densely 
punctate,  punctures  moderate  and  just  a  little  larger  than  those  on  the 
head ;  apical  angles  narrowly  rounded ;  basal  angles  rectangular. 

Propleura  densely  punctate. 

Elytra  widest  at  the  posterior  third,  a  little  more  than  one-half  longer 
than  wide ;  humeri  small ;  sides  diverging  and  feebly  arcuate  to  apical  half, 
thence  more  strongly  and  broadly  arcuate  to  apex,  the  latter  obtusely 
rounded ;  disk  rather  strongly  convex  posteriorly,  less  so  anteriorly,  evenly 
and  arcuately  declivous  posteriorly,  densely  ruguloso-punctate,  striate, 
striae  fine  and  broken  up  into  short  lines  by  the  rugulae,  sometimes  scarcely 
evident. 

Male. — Form  somewhat  narrower.     Anterior  tarsi  moderately  dilated. 

Measurements. — Length,  12-20  mm. ;  width,  5.2-7.75  mm. 

Habitat. — Under  bark. 

Holotype,  female,  and  allotype,  male,  in  my  own  collection.  Para- 
types  in  my  own  and  Mr.  F.  W.  Nunenmacher's  collection. 

Type  locality. — Male  taken  at  Sisson,  and  the  female  at  Shasta 
Retreat,  Siskiyou  County,  California. 

Distribution. — California  (Humboldt  County:  Green  Point  Ranch, 
elevation  1500  feet,  June;  Siskiyou  County:  Sisson,  July;  Shasta  Retreat, 
elevation  2416  feet,  July;  Shasta  County,  May). 

Remarks. — Simulator  appears  to  be  absolutely  distinct  from  opacus 
Lee.  The  latter  is  apterous  and  the  pronotum  is  much  more  strongly 
convex.  They  may  or  may  not  inhabit  the  same  territory.  Opacus  is 
taken  quite  commonly  in  Calaveras  County.  As  Horn  states,  the  "winged 
species  has  the  elytra  depressed,  more  elongate  and  less  rounded  on  the 
sides ;  the  thorax  is  also  less  convex,  rather  broader  and  with  less  rounded 
sides."  These  remarks  apply  to  simulator,  which  Dr.  Horn  considered  a 
form  of  opacus. 

Helops  regulus,  new  species. — Form  elongate  oval,  slightly  narrowed 
anteriorly,  convex.  Color  nigro-piceous,  scarcely  submetallic,  shining; 
legs  and  antennae  more  or  less  rufo-piceous ;  beneath  rather  rufo-piceous. 

Head  convex,  more  or  less  deeply  and  transversely  impressed  behind 
the  frontal  suture,  rather  coarsely,  strongly  and  moderately  densely  punc- 


228      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

tate,  punctures  rounded,  scarcely  coalescent.  Antennae  moderately  short 
and  somewhat  stout,  reaching  a  little  beyond  the  base  of  the  pronotum. 

Pronotum  nearly  as  long  as  wide,  not  noticeably  transverse,  appar- 
ently widest  at  the  middle ;  sides  rather  moderately  arcuate,  finely  beaded ; 
apex  and  base  subtruncate,  not  strongly  beaded;  apex  a  little  narrower 
than  base;  apical  angles  subrectangular ;  basal  angles  obtuse  and  some- 
what rounded ;  disk  moderately  strongly  convex  from  side  to  side,  quite 
narrowly  explanate  and  impressed  within  the  bead;  punctate,  punctures 
more  or  less  irregularly  placed,  moderately  coarse,  partly  somewhat 
slightly  elongate  at  the  sides. 

Elytra  oval,  about  three-sixths  longer  than  wide ;  sides  evenly  arcuate 
and  subparallel,  obtusely  rounded  at  apex;  base  truncate,  humeri  small 
and  slightly  prominent  anteriorly  and  distinct;  disk  rather  evenly  convex, 
striae  of  small  punctures  which  are  slightly  irregularly  spaced  and  feebly 
impressed;  intervals  sometimes  slightly  convex,  finely  and  sparsely  punc- 
tulate,  not  tuberculate  nor  rugulose  on  the  apical  declivity. 

Propleura  finely  and  more  or  less  irregularly  longitudinally  rugulose. 

Abdomen  somewhat  rugulose  and  rather  more  thickly  punctate  on  the 
first  two  than  on  the  succeeding  segments. 

Tarsi  clothed  beneath  with  rather  soft  yellow  hairs. 

Male. — Rather  narrower,  antennae  slightly  stouter. 

Female. — Somewhat  broader,  antennae  somewhat  more  slender. 

Measurements.-. — Female  holotype  :    Length,  5.8  mm. ;  width,  2.2  mm. 

The  accompanying  male  allotype  measures  7.5  mm.  in  length ;  the 
elytra  are  separated  and  the  wings  are  rudimentary. 

The  types  are  in  my  own  collection.  Collected  by  Mr.  W.  M.  Mann, 
on  May  15,  1904.  About  twenty  specimens  in  various  collections  have 
been  studied. 

Type  locality. — Wawawai,  State  of  Washington. 

Regulus  was  first  referred  to  pernitens  Lee.  with  considerable  doubt. 
It  at  first  sight  resembles  aereus  Germ,  in  general  facies,  from  which  it 
differs  in  the  narrower  elytra  and  different  punctuation  of  the  head  and 
pronotum.  In  pernitens  the  pronotum  is  distinctly  transverse,  antennae 
longer  and  more  slender,  elytra  broader,  sides  of  the  pronotum  distinctly 
explanate  with  margin  reflexed.  In  both  species  the  mentum  is  longi- 
tudinally carinate  at  middle  with  the  sides  opaque,  carina  smooth  and 
shining.  In  regulus  the  pronotum  is  rather  more  punctate  than  rugulose, 
the  reverse  in  pernitens.  The  propleura  in  aereus  is  sculptured  with 
rather  regular,  strong,  parallel,  longitudinal  rugae,  some  of  which  divide. 

Helops  obtusangula,  new  species. — Form  elongate  ovate,  subparallel, 
moderately  depressed.  Color  brownish  to  piceous,  not  metallic ;  legs  and 
under  surface  more  or  less  rufo-piceous.  Apterous. 


HELOPS  OBTUSANGULA  229 

Head  subquadrate,  front  scarcely  convex  between  the  eyes,  flattened 
toward  the  epistoma,  sides  of  front  moderately  prominent;  feebly  and 
transversely  constricted  behind  the  eyes;  densely,  not  coarsely  punctate, 
punctures  more  or  less  slightly,  coalescent.  Eyes  slightly  prominent. 
Antennae  slender,  outer  joints  slightly  compressed,  last  three  just  a  little 
wider,  eleventh  oval. 

Pronotum  subquadrate,  about  a  fourth  wider  than  long ;  apex  almost 
truncate,  somewhat  feebly  arcuate  centrally ;  apical  angles  obtuse ;  sides 
feebly  arcuate  from  apex  to  base,  most  strongly  so  anteriorly,  less  so 
posteriorly  and  just  the  least  converging,  marginal  bead  not  strong  or 
prominent;  base  almost  truncate;  basal  angles  obtuse  and  narrowly 
rounded,  scarcely  distinct ;  disk  not  strongly  convex,  most  so  anteriorly 
and  centrally  at  apex,  there  rather  strongly  declivous  laterally,  vaguely 
and  transversely  impressed  in  front  of  base,  surface  very  slightly  flattened 
in  lateral  fourths,  densely  punctate,  punctures  small,  more  or  less 
coalescent  longitudinally,  intervals  not  conspicuous. 

Pro  pleura  distinctly  and  not  coarsely  longitudinally  rugulose ;  rugulae 
uniting  to  a  greater  or  less  extent,  with  large,  shallow  and  scarcely  distinct 
punctures  in  the  broad  sulci  between. 

Elytra  elongate  oval,  a  little  more  than  twice  as  long  as  wide,  moder- 
ately evenly  convex  from  side  to  side,  a  little  wider  than  the  pronotum, 
humeri  obtusely  rounded,  base  transverse ;  surface  finely  striate,  striae  not 
noticeably  punctate,  intervals  finely,  irregularly  and  somewhat  sparsely 
punctate,  feebly  convex  especially  on  apical  declivity,  there  with  feeble, 
small  tuberculiform  eminences ;  sides  parallel,  feebly  arcuate,  apex  some- 
what ogival  and  rather  narrowly  rounded. 

Abdomen  shining,  rather  evenly  and  not  densely  nor  coarsely  punc- 
tate. Legs  rather  short. 

Male. — Narrower.  Antennae  elongate,  joints  eight  to  eleven  inclu- 
sive at  least  twice  as  long  as  wide,  not  incrassate,  scarcely  compressed. 
Anterior  tarsi  feebly  dilated,  and  with  the  middle  tarsi  quite  densely 
clothed  beneath  with  yellow  hairs. 

Female. — Somewhat  broader.  Antennae  shorter,  outer  four  joints 
distinctly  broader,  not  twice  as  long  as  wide,  ninth  and  tenth  subtriangular, 
slightly  compressed.  Anterior  tarsi  very  slightly  dilated.  All  the  tarsi 
clothed  beneath  with  yellow  hairs,  denser  on  the  anterior  and  middle  tarsi. 

Measurements. — Length   (Types),  10.5-S  mm.;  width,  3.5-2.6  mm. 

Holotype,  female,  larger  sex,  in  the  collection  of  the  California 
Academy  of  Sciences.  Allotype,  male,  and  paratypes  in  the  collection  of 
the  author. 

Holotype  collected  by  Mr.  Geo.  R.  Wilson ;  allotype  and  paratypes  by 
Dr.  E.  C.  Van  Dyke. 


230      NEW  SPECIES  MELYRIDAE,  CHRYSOMELIDAE  AND  TENEBRIONIDAE 

Type  locality. — Corona,  California  (Holotype)  ;  Los  Angeles,  Cali- 
fornia (Allotype). 

Obtusangula  has  been  labeled  as  angustus  Lee.  in  my  collection  for 
many  years.  An  examination  of  my  series  shows  it  to  be  wingless  and 
the  propleura  distinctly  but  not  strongly  rugulose,  punctures  not  evident. 

Horn  states  that  angustus  is  winged  and  that  the  propleura  are 
coarsely  punctate. 

Obtusangula  is  related  to  stenotrichoides  Blais.,  similar  in  form,  but 
less  elongate,  less  robust,  and  less  convex,  integuments  thinner  and  the 
small  tubercles  of  the  lateral  and  apical  intervals  of  the  elytra  are  less 
evident,  besides,  the  basal  angles  of  the  pronotum  are  obtuse  and  more  or 
less  slightly  rounded. 

In  stenotrichoides  the  basal  angles  are  distinct,  propleura  punctato- 
rugulose,  punctures  distinct  and  shallow  on  the  coxal  convexities ;  outer 
joints  of  the  antennae  slightly  compressed  in  the  male  and  twice  as  long 
as  wide;  antennae  more  elongate  in  the  female  than  in  the  same  sex  of 
Obtusangula;  with  joints  six,  seven  and  eight  rather  more  elongate.  In 
strigicollis  Horn  the  form  is  narrower,  elytra  scarcely  wider  than  the 
pronotum,  the  latter  is  distinctly  longitudinally  strigose,  antennae  heavier, 
propleura  distinctly  punctate.  Strigicollis  and  obtusangula  were  taken  at 
the  same  time  at  Corona,  California,  by  Mr.  Wilson. 

Attenitatus  belongs  to  the  same  group  of  species.  My  only  specimen 
referred  to  this  species  was  collected  by  Mr.  Nunenmacher  at  Goldfield, 
Nevada.  The  form  is  almost  subcylindrical,  pronotum  almost  discretely 
and  rather  sparsely  punctate,  antennae  with  joints  four  to  eleven  subequal, 
propleura  coarsely  punctate,  each  puncture  with  a  small  yellow  hair.  I 
can  not  agree  with  Horn  in  considering  that  attenuatus  should  be  asso- 
ciated with  bachei  Lee.  It  is  very  desirable  to  study  the  genitalia  of  the 
different  species,  but  the  scarcity  of  specimens  forbids  any  dissection  at 
the  present  time. 

The  above  species,  as  I  view  them,  may  be  separated  as  follows : 

Body  winged  augustus  Lee. 

Body  wingless  - _ _.     1 

1.  Pronotum  distinctly  strigose -strigicollis  Horn. 

-.     Pronotum  distinctly  punctate  , 2 

2.  Body  distinctly  subcylindrical attenuatus  Lee. 

-.     Body  more  or  less  depressed 3 

3.  Antennae  elongate  in  both  sexes,  slightly  longer  in  the  male ;  basal  angles  of 

the  pronotum  distinct stenotrichoides  Blais. 

-.  Antennae  elongate  in  the  male,  distinctly  shorter  in  the  female ;  basal  angles 

obtuse  and  slightly  rounded _ obtusangula,  n.  sp. 

In  the  above  species  the  color  is  brownish  to  piceous,  legs  more  or 
less  rufous  with  the  abdomen  darker  rufo-piceous. 


ADDENDUM  231 

ADDENDUM 

Mr.  Ralph  Hopping,  of  the  Canadian  Forestry  Service,  has  very 
recently  submitted  to  me  a  species  of  Listrus,  collected  at  Vernon,  B.  C. 
It  has  proved  to  be  an  undescribed  species  to  which  I  have  given  the 
name  provincialis,  n.  sp.  The  description  will  in  all  probability  appear  in 
the  Canadian  Entomologist  in  the  near  future.  The  types  will  be  placed 
in  the  National  Collection  at  Ottawa.  Provincialis  should  follow  difficilis 
Lee.  in  the  synoptical  arrangement  given  above. 


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